^Pounds, J.A. and Crump, M.L. 1994. Amphibian declines and climate disturbance: The case of the golden toad and the harlequin frog. Conservation Biology8: Page 72-85
^Pounds, J.A., and Crump, M.L. 1987. "Harlequin Frogs Along a Tropical Montane Stream: Aggregation and the Risk of Predation by Frog-Eating Flies." Biotropica19(4): Page 306-309.
^Crump, M.L. 1986. Homing and site fidelity in a Neotropical frog, Atelopus varius (Bufonidae). Copeia 1986(4): 1007-1009
^Lips, K.R. 1999. Mass mortality and population declines of anurans at an upland site in western Panama. Conservation Biology13: Page 117-125
^Lips, K.R., Green, D.E. and Papendick, R. 2003. Chytridiomycosis in wild frogs from southern Costa Rica. Journal of Herpetology37: page. 215-218