This list of fossilreptiles described in 2020 is a list of new taxa of fossil reptiles that were described during the year 2020, as well as other significant discoveries and events related to reptile paleontology that occurred in 2020.
New fossil material of squamates is described from the Upper Cretaceous Fruitland and Kirtland formations (New Mexico, United States) by Woolley, Smith & Sertich (2020), expanding known taxonomic and morphological diversity of lizards within the "Hunter Wash Local Fauna".[14]
A study on the affinities of putative gekkotan eggshells from the Late Cretaceous of Europe is published by Choi et al. (2020), who interpret the fossil material of Pseudogeckoolithus as theropod eggshells.[15]
A dentary of a cnemidophorine teiid is described from the Miocene of the Ogallala Group (Nebraska, United States) by Scarpetta (2020), who evaluates the implications of this specimen for the knowledge of the evolutionary history of cnemidophorines in North America during the Neogene.[17]
The first known fossil example of an iguana nesting burrow is reported from the Pleistocene Grotto Beach Formation (The Bahamas) by Martin et al. (2020).[18]
A study on the anatomy of the skull of Ophisaurus acuminatus, and on the taxonomic validity of this species, is published by Klembara & Čerňanský (2020).[19]
Fossil material of the monitor lizards is reported for the first time from the late Miocene localities in Armenia and Georgia by Vasilyan & Bukhsianidze (2020).[20]
A study on the evolutionary history of mosasauroids, comparing their evolutionary rates and traits to those of plesiosaurs and aiming to determine whether the rise and diversification of mosasauroids was influenced by competition with or disappearance of some plesiosaur taxa, is published by Madzia & Cau (2020).[21]
Grigoriev & Grabovskiy (2020) describe new fossil material of a tylosaurine from the Upper Cretaceous (Turonian) of the Chukotka Region (Russia), representing one of the oldest and northernmost mosasaur records reported so far, and evaluate the implications of this fossil material (as well as mosasaur fossils from the Santonian of the Komi Republic and from the Campanian–Maastrichtian of the Sakhalin Island) for the knowledge of the paleogeography and possible migrations of Arctic mosasaurs.[22]
A study on pathological features of a specimen of Prognathodon (belonging or related to the species P. sectorius) from the MaastrichtianGulpen Formation (the Netherlands) is published by Bastiaans et al. (2020), who consider it most likely that this specimen was bitten in the snout by a large, possibly conspecific mosasaur, making it one of the few specimens with unambiguous evidence of agonistic interactions amongst mosasaurs.[23]
A study on the morphology of the snout of Taniwhasaurus antarcticus, indicating the presence of a complex internal neurovascular system of branched channels similar to systems present in extant aquatic vertebrates such as cetaceans and crocodiles, is published by Álvarez–Herrera, Agnolin & Novas (2020).[24]
Zietlow (2020) recovers growth series of Tylosaurus proriger and T. nepaeolicus, and tests the hypothesis that T. kansasensis represent juveniles of T. nepaeolicus.[25]
Redescription of Palaeophis oweni is published by Georgalis, Del Favero & Delfino (2020).[26]
Eocene snake vertebrae from Landana and Sassa-Zao (Angola), originally referred to Palaeophis aff. typhaeus, are assigned to the species Palaeophis africanus by Folie et al. (2020), who interpret these fossils as confirming the aquatic capabilities of palaeophiid snakes, and evaluate the implications of these fossils for the debate on the existence of primitive and advanced grades among palaeophiid snakes.[27]
New information on the anatomy of the Eocene fossil boas Messelophis variatus and Rieppelophis ermannorum, based on data from new specimens from the Messel pit (Germany), is presented by Scanferla & Smith (2020).[28]
Description of new fossil material of Thaumastophis missiaeni from the Eocene Cambay Shale (India) and a study on the phylogenetic relationships of this snake is published by Zaher et al. (2020), who name a new family Thaumastophiidae.[29]
New anatomical features of the holotype specimen of Cartorhynchus lenticarpus revealed by CT scanning, including unique dentition, are reported by Huang et al. (2020), who evaluate the implications of this species for the knowledge of the evolution of tooth morphology and diet in basalichthyosauriforms.[36]
A caudal vertebra of a basal ichthyosauriform similar to grippidians, representing the youngest record of basal ichthyosauriforms to date, is described from the Middle Triassic (Ladinian) of Mallorca (Spain) by Matamales-Andreu et al. (2020).[37]
A study on the age of the fossils of Thaisaurus chonglakmanii is published by Tongtherm et al. (2020).[38]
A study on the tempo and mode of the morphological evolution of ichthyosaurs is published by Moon & Stubbs (2020).[39]
A study on skeletal pathologies in ichthyosaur specimens from the Middle TriassicBesano Formation and the Lower JurassicPosidonia Shale, evaluating their implications for the knowledge of changing locomotory and behavioural constraints affecting different ichthyosaur taxa through time, is published by Pardo-Pérez, Kear & Maxwell (2020).[40]
Two new specimens of Mixosaurus cornalianus, preserving evidence of the presence of a dorsal fin and a well-developed, triangular dorsal lobe of the caudal fin in this species, are described from the AnisianBesano Formation (Italy) by Renesto et al. (2020).[41]
Jiang et al. (2020) report the discovery of remains of a thalattosaur belonging to the genus Xinpusaurus in the abdominal region of a specimen of Guizhouichthyosaurus from the Middle Triassic (Ladinian) Zhuganpo Member of the Falang Formation (China), and interpret this finding as likely the oldest evidence for predation on megafauna by marine tetrapods reported so far.[42]
A study on the anatomy of the holotype specimen of Temnodontosaurus crassimanus is published by Swaby & Lomax (2020), who consider T. crassimanus to be a valid species.[43]
Description of the most complete and best-preserved skeleton of Suevoleviathan integer is published by Maisch (2020).[44]
Partial skeleton of an ophthalmosaurid ichthyosaur, found with an ichthyosaur tooth (probably not belonging to the same specimen) stuck on its rib, is described from the Upper Jurassic Rosso Ammonitico Veronese Formation (Italy) by Serafini et al. (2020), possibly representing the first evidence of scavenging between two ichthyosaurs reported so far.[45]
A study on the anatomy and phylogenetic relationships of Maiaspondylus lindoei, "Ophthalmosaurus" cantabrigiensis and "Platypterygius" ochevi is published by Zverkov & Grigoriev (2020), who transfer "O". cantabrigiensis to the genus Maiaspondylus, and consider "P. ochevi to be a junior synonym of M. cantabrigiensis.[46]
Description of a new specimen of Muiscasaurus catheti from the AptianPaja Formation (Colombia), providing new information on the anatomy of this taxon, and a study on the phylogenetic relationships of M. catheti is published by Páramo-Fonsecaet al. (2020).[47]
A study on the osteology and evolution of the temporal region of the skull of placodonts is published by Maisch et al. (2020).[52]
A study on the anatomy of the skull and braincase of Parahenodus atancensis, and on the anatomy of the reconstructed brain, inner ear and neurosensory structures of this taxon, is published by De Miguel Chaves et al. (2020).[53]
New fossil material of Lariosaurus sanxiaensis is described from the Lower TriassicJialingjiang Formation (China) by Li & Liu (2020), who also study the phylogenetic relationships of this taxon, as well as the predator-prey relationship in the associated fauna and their implications for the knowledge of the biotic recovery after the Permian–Triassic extinction event.[54]
New fossil material of cryptoclidid plesiosaurs, including the first occurrence of Vinialesaurus in the Southern Hemisphere, is described from the Jurassic of the Atacama Desert by Otero et al. (2020).[55]
Redescription of the holotype specimen of Aphrosaurus furlongi and a study on the evolution and phylogenetic relationships of elasmosaurid plesiosaurs is published by O'Gorman (2020), who names a new clade Euelasmosaurida.[56]
The first non-aristonectine elasmosaurid skeleton from Antarctica that preserves an associated lower jaw is described by O'Gorman et al. (2020).[57]
An isolated cervicalcentrum of a brachaucheninepliosaurid is described from the Cenomanian of Russia by Zverkov & Pervushov (2020), who interpret this fossil as belonging to one of the largest known pliosaurids, and consider it to be evidence of survival gigantic pliosaurids into the Cenomanian.[58]
A study on the evolution of the short-necked plesiosaurs throughout the Jurassic and Cretaceous periods is published by Fischer et al. (2020).[59]
A study on the evolution of turtle skull architecture, aiming to assess the functional significance of changes in their skull architecture during feeding on the basis of data from extant and fossil taxa, is published by Ferreira et al. (2020).[84]
A study on the relation between ecology and shell shape in extant turtles, and its implications for the knowledge of the ecology of fossil turtles, is published by Dziomber, Joyce & Foth (2020).[85]
A study on the early evolution of turtles during the Triassic period is published by de la Fuente, Sterli & Krapovickas (2020).[86]
A study on non-marine turtle distribution and diversity from the Late Triassic to the Paleogene is published by Cleary et al. (2020).[87]
Szczygielski (2020) revises the first described Triassic turtle Chelytherium obscurum, and considers it to be synonymous with Proterochersis robusta.[88]
Description of new fossil material of Indochelys spatulata from the JurassicKota Formation (India), and a study on the anatomy and phylogenetic relationships of this taxon, is published by Joyce & Bandyopadhyay (2020).[89]
The first three-dimensional reconstruction of the skull of Kallokibotion bajazidi is presented by Martín-Jiménez, Codrea & Pérez-García (2020).[90]
Fossil material of an indeterminate member of Pelomedusoides is described from the ValanginianRosablanca Formation (Colombia) by Cadena (2020), who interprets this finding as additional evidence supporting the occurrence of Pelomedusoides during the Valanginian in northern South America.[91]
New specimens of Araripemys barretoi, providing new information on the morphological variation within this species, are described from the Crato Formation and Romualdo Formation (Brazil) by Limaverde et al. (2020).[92]
Cadena et al. (2020) describe new fossil material of Stupendemys geographicus from the Miocene of Venezuela and Colombia, providing new information on the anatomy and paleobiology of this species.[95]
Description of the anatomy of the skull of Pleurosternon bullockii is published by Evers, Rollot & Joyce (2020).[97]
Redescription of the anatomy of the skull and mandible of Sandownia harrisi is published by Evers & Joyce (2020).[98]
New information on the anatomy and stratigraphic and geographic distribution of Anosteira pulchra is presented by Adrian et al. (2020).[99]
A revision of the extinct geoemydidEchmatemys from North America, based mainly on data from a slab containing several turtle shells collected from the Bridgerian of Levett Creek (Wyoming, United States), is published by Vlachos (2020).[100]
A study on the phylogenetic relationships of a putative testudinoidCardichelyon rogerwoodi is published by Joyce & Claude (2020), who consider it more likely that this taxon is a member of Kinosternoidea.[101]
A member of the family Doswelliidae; a new genus for "Doswellia" sixmilensis Heckert, Lucas & Spielmann (2012).
Research
New Early Triassic archosauriform track assemblage is described from the Gardetta Plateau (Western Alps, Italy) by Petti et al. (2020), who interpret this finding as evidence of the presence of archosauriforms at low latitudes soon after the Permian–Triassic extinction event, and name a new ichnotaxon Isochirotherium gardettensis.[105]
Postcranial material of an erythrosuchid from the Rassypnaya locality (Olenekian; Orenburg Oblast, Russia), previously referred to Vjushkovia triplicostata, is referred to the species Garjainia prima by Maidmentet al. (2020), who evalue the implications of this fossil material for the knowledge of the phylogenetic relationships, body mass and locomotor musculature of erythrosuchids.[106]
Redescription of the anatomy of the holotype specimen of Chanaresuchus bonapartei and a study on the phylogenetic relationships of this species is published by Trotteyn & Ezcurra (2020).[107]
Redescription of the anatomy of the skull and mandible of Euparkeria capensis is published Sookias et al. (2020).[108]
A study on the joint mobility of the hindlimb of Euparkeria capensis, and on its implications for the knowledge of the evolution of the locomotor capabilities of archosaurs, is published by Demuth, Rayfield & Hutchinson (2020).[109]
An assemblage of at least 21 phytosaur specimens dominated by juveniles and subadults is described from the Upper Triassic Tiki Formation (India) by Datta, Mukherjee & Ray (2020), who interpret this finding as likely evidence of parental care in phytosaurs, and study the taphonomy of the assemblage.[111]
A study on the evolution of the skull shape in phytosaurs is published by Datta, Sharma & Ray (2020).[112]
A study comparing teeth microwear textures in Machaeroprosopus pristinus, Mystriosuchus planirostris, Nicrosaurus kapffi, N. meyeri and "Smilosuchus" lithodendrorum, aiming to determine whether microwear texture differences reflect dietary differences between phytosaur species, is published by Bestwick et al. (2020).[113]
A member of the family Acleistorhinidae. The type species is C. lundi. Announced in 2019;[114] the correction including the required ZooBank accession number was published in 2020.[115]
A diapsid of uncertain phylogenetic placement. Originally described as a member of Avialae[123] but subsequently argued to be a lizard.[124] Genus includes new species O. khaungraae. Its status as a validly named taxon is controversial because the scientific article it was named in was subsequently retracted.[125][126]
A procolophonid; a new genus for "Eumetabolodon" dongshengensis Li (1983).
Research
A study on fracture planes (unossified regions in the middle of vertebral centra) in tail vertebrae of mesosaurs is published by MacDougall et al. (2020), who argue that mesosaurs were theoretically capable of tail autotomy, but probably did not utilize this ability.[134]
A study on patterns of tooth development and replacement in Belebey and Bolosaurus, indicating that bolosaurid teeth had thecodont implantation with deep roots, is published by Snyder et al. (2020).[135]
Revision of Pachypes-like footprints from the Cisuralian–Guadalupian of Europe and North America is published by Marchetti et al. (2020), who date the earliest known occurrence of Pachypes to the Artinskian, interpret the footprints belonging to the ichnospecies Pachypes ollieri as produced by nycteroleterpareiasauromorphs, and argue that the earliest occurrences of pareiasauromorph footprints precede the earliest occurrence of this group in the skeletal record by at least 10 million years.[136]
A study on the dental wear along the tooth rows of nearly one hundred jaws of Captorhinus aguti, indicating that this reptile preferred to feed using the right side of the jaw, is published by Reiszet al. (2020).[137]
Redescription of Cargninia enigmatica is published by Vivar et al. (2020).[138]
A study on the phylogenetic relationships and evolutionary history of sphenodontian reptiles is published by Simões, Caldwell & Pierce (2020).[139]
A study on the feeding mechanics and ecology of Clevosaurus hudsoni and C. cambrica, as indicated by their bite force, resistance of skull bones to bending and torsion, and the distribution of stresses in the jaws during biting, is published by Chambi-Trowell et al. (2020).[140]
A study on the anatomy and phylogenetic relationships of Colobops noviportensis is published by Scheyer et al. (2020), who reinterpret this taxon as a probable rhynchocephalian.[141]
A study on the morphology of teeth of Priosphenodon avelasi is published by LeBlanc et al. (2020).[142]
Partial skeleton of a small reptile, probably a juvenile specimen of Eusaurosphargis dalsassoi, is described from the AnisianBuchenstein Formation (northern Dolomites, Italy) by Renesto, Kustatscher & Gianolla (2020), who interpret this finding as possible evidence of that the lands emerged near the basins of the northern Dolomites, Besano Formation and Prosanto Formation had a similar reptilian fauna during the middle-late Anisian.[143]
A study on the anatomy of the skull of Champsosaurus lindoei is published by Dudgeon et al. (2020), who evaluate the morphology of a putative neomorphic bone in the skull and its possible developmental and functional origins.[145]
A study on the internal anatomy of the skull of Champsosaurus lindoei and C. natator, and on their probable sensory abilities, is published by Dudgeon et al. (2020).[146]
A study on the Triassic fossil record and evolution of non-archosaurian archosauromorph reptiles in South America is published by Ezcurra et al. (2020), who also identify the first record of the family Proterosuchidae from South America (partial braincase from the Buena Vista Formation, Uruguay).[147]
New fossil material of tanystropheid and azendohsaurid archosauromorphs, providing new information on the diversity of Late Triassic archosauromorph reptiles in North America, is described from the Lamy Quarry south of the town of Lamy (Garita Creek Formation; New Mexico, United States) by Hégron et al. (2020).[148]
Redescription of the anatomy of the skeleton of Macrocnemus fuyuanensis is published by Scheyer et al. (2020).[149]
Description of the morphology of the skull of Macrocnemus bassanii is published by Miedema et al. (2020).[150]
A study on the morphology of the skull of Tanystropheus hydroides is published by Spiekman et al. (2020).[151]
A study on the dynamics of phenotypic and molecular evolution of reptiles during the early diversification of the major lineages of diapsid reptiles in the Permian and Triassic periods, and during the evolution of lepidosaurs from the Jurassic to the present, is published by Simões et al. (2020).[153]
A large, soft-shelled egg, most closely resembling eggs of extant lizards and snakes and possibly produced by a mosasaur, is described from the Upper Cretaceous Lopez de Bertodano Formation (Antarctica) by Legendre et al. (2020), who name a new ootaxonAntarcticoolithus bradyi.[154]
A study on the ecological diversity of Mesozoic marine tetrapods is published by Reeves et al. (2020).[155]
A study on the evolution of the archosauromorph ankle, aiming to test the hypothesis of fusion between the centrale and astragalus and the alternative hypothesis of a complete loss of the centrale, based on embryological and palaeontological data, is published by Blanco, Ezcurra & Bona (2020).[157]
^Catherine R. C. Strong; Michael W. Caldwell; Takuya Konishi; Alessandro Palci (2020). "A new species of longirostrine plioplatecarpine mosasaur (Squamata: Mosasauridae) from the Late Cretaceous of Morocco, with a re-evaluation of the problematic taxon 'Platecarpus' ptychodon". Journal of Systematic Palaeontology. 18 (21): 1769–1804. Bibcode:2020JSPal..18.1769S. doi:10.1080/14772019.2020.1818322. S2CID224978215.
^Annelise Folie; Florias Mees; Thierry De Putter; Thierry Smith (2020). "Presence of the large aquatic snake Palaeophis africanus in the middle Eocene marine margin of the Congo Basin, Cabinda, Angola". Geobios. 66–67: 45–54. doi:10.1016/j.geobios.2020.11.002. S2CID229411514.
^Agustín Scanferla; Krister T. Smith (2020). "Additional anatomical information on the Eocene minute boas Messelophis variatus and Rieppelophis ermannorum (Messel Formation, Germany)". Vertebrate Zoology. 70 (4): 615–620. doi:10.26049/VZ70-4-2020-06.
^Hussam Zaher; Annelise Folie; Ana B. Quadros; Rajendra S. Rana; Kishor Kumar; Kenneth D. Rose; Mohamed Fahmy; Thierry Smith (2020). "Additional vertebral material of Thaumastophis (Serpentes: Caenophidia) from the early Eocene of India provides new insights on the early diversification of colubroidean snakes". Geobios. 66–67: 35–43. doi:10.1016/j.geobios.2020.06.009. S2CID225415393.
^Jair Israel Barrientos Lara; Jesús Alvarado Ortega; Marta S.Fernández (2020). "Acuetzpalin carranzai gen et sp. nov. A new ophthalmosauridae (Ichthyosauria) from the Upper Jurassic of Durango, North Mexico". Journal of South American Earth Sciences. 98: Article 102456. Bibcode:2020JSAES..9802456B. doi:10.1016/j.jsames.2019.102456. S2CID213102661.
^Nicole Klein; Lars Schmitz; Tanja Wintrich; P. Martin Sander (2020). "A new cymbospondylid ichthyosaur (Ichthyosauria) from the Middle Triassic (Anisian) of the Augusta Mountains, Nevada, USA". Journal of Systematic Palaeontology. 18 (14): 1167–1191. Bibcode:2020JSPal..18.1167K. doi:10.1080/14772019.2020.1748132. S2CID219078178.
^Nikolay G. Zverkov; Megan L. Jacobs (2020). "Revision of Nannopterygius (Ichthyosauria: Ophthalmosauridae): reappraisal of the 'inaccessible' holotype resolves a taxonomic tangle and reveals an obscure ophthalmosaurid lineage with a wide distribution". Zoological Journal of the Linnean Society. 191 (1): 228–275. doi:10.1093/zoolinnean/zlaa028.
^Yu Qiao; Masaya Iijima; Jun Liu (2020). "The largest hupehsuchian (Reptilia, Ichthyosauromorpha) from the Lower Triassic of South China indicates early establishment of high predation pressure after the Permo-Triassic mass extinction". Journal of Vertebrate Paleontology. 39 (5): e1719122. doi:10.1080/02724634.2019.1719122. S2CID215756095.
^Michael W. Maisch (2020). "The best-preserved skeleton of Suevoleviathan integer (Bronn, 1844) (Reptilia: Ichthyosauria) from the Lower Jurassic of south-western Germany, with a discussion of the genus". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 297 (2): 153–172. doi:10.1127/njgpa/2020/0918. S2CID225493146.
^Giovanni Serafini; Jacopo Amalfitano; Miriam Cobianchi; Beatrice Fornaciari; Erin E. Maxwell; Cesare Andrea Papazzoni; Guido Roghi; Luca Giusberti (2020). "Evidence of opportunistic feeding between ichthyosaurs and the oldest occurrence of the hexanchid shark Notidanodon from the Upper Jurassic of Northern Italy". Rivista Italiana di Paleontologia e Stratigrafia. 126 (3): 629–655. doi:10.13130/2039-4942/14078. S2CID229243429.
^Nikolay G. Zverkov; Dmitry V. Grigoriev (2020). "An unrevealed lineage of platypterygiines (Ichthyosauria) with peculiar forefin structure and semiglobal distribution in the mid-Cretaceous (Albian–Cenomanian)". Cretaceous Research. 115: Article 104550. Bibcode:2020CrRes.11504550Z. doi:10.1016/j.cretres.2020.104550. S2CID225721312.
^María Eurídice Páramo-Fonseca; Javier García Guerrero; Cristian David Benavides-Cabra; Santiago Padilla Bernal; Antonio José Castañeda-Gómez (2020). "A benchmark specimen of Muiscasaurus catheti from the upper Aptian of Villa de Leiva, Colombia: new anatomical features and phylogenetic implications". Cretaceous Research. 119: Article 104685. doi:10.1016/j.cretres.2020.104685. S2CID228965354.
^Rodrigo A. Otero; Sergio Soto-Acuña (2020). "Wunyelfia maulensis gen. et sp. nov., a new basal aristonectine (Plesiosauria, Elasmosauridae) from the Upper Cretaceous of central Chile". Cretaceous Research. 118: Article 104651. doi:10.1016/j.cretres.2020.104651. S2CID224975253.
^José Patricio O'Gorman; Paula Bona; Martín de los Reyes; Maria Eugenia Raffi; Marcelo Reguero (2020). "A non-aristonectine plesiosaur from Antarctica reveals new data on the mandibular symphysis of elasmosaurids". Alcheringa: An Australasian Journal of Palaeontology. 44 (4): 565–576. Bibcode:2020Alch...44..565O. doi:10.1080/03115518.2020.1824261. S2CID229399482.
^A. Pérez-García (2020). "A new bothremydid turtle (Pleurodira) from the Upper Cretaceous (Cenomanian) of Madagascar". Cretaceous Research. 118: Article 104645. doi:10.1016/j.cretres.2020.104645. S2CID225033660.
^Adán Pérez-García; Evangelos Vlachos; Xabier Murelaga (2020). "A large testudinid with African affinities in the post-Messinian (lower Pliocene) record of south-eastern Spain". Palaeontology. 63 (3): 497–512. doi:10.1111/pala.12468. S2CID214232312.
^Gerardo Carbot-Chanona; Gustavo Rivera-Velázquez; Eduardo Jiménez-Hidalgo; Víctor Hugo Reynoso (2020). "The first Pan-Carettochelys turtle in the Neogene of the American continent and its paleobiogeographical relevance". Journal of South American Earth Sciences. 104: Article 102925. Bibcode:2020JSAES.10402925C. doi:10.1016/j.jsames.2020.102925. S2CID224976641.
^Adán Pérez-García; Thierry Smith (2020). "Systematics and diversity of the giant soft-shelled turtles (Cryptodira, Trionychidae) from the earliest Eocene of Belgium". Geobios. 66–67: 15–34. doi:10.1016/j.geobios.2020.07.006. S2CID229426614.
^Oleg Redkozubov; Elena Syromyatnikova; Anna Gnetneva; Igor Danilov (2020). "First reliable records of the genus Chersine (Testudines: Testudinidae) from the Pliocene of Moldova". Historical Biology: An International Journal of Paleobiology. 33 (11): 3046–3055. doi:10.1080/08912963.2020.1846531. S2CID229407378.
^Diogo Lins Batista; Ismar de Souza Carvalho; Marcelo S. de la Fuente (2020). "A new Cretaceous (Pleurodira:Pelomedusoides) from the Lower Cretaceous of Parnaíba Basin, Brazil". Journal of South American Earth Sciences. 105: Article 102872. doi:10.1016/j.jsames.2020.102872. S2CID224959263.
^Edwin-Alberto Cadena; Andrés Vanegas; Carlos Jaramillo; John M. Cottle; Thomas A. Johnson (2020). "A new Miocene turtle from Colombia sheds light on the evolutionary history of the extant genus Mesoclemmys Gray, 1873". Journal of Vertebrate Paleontology. 39 (5): e1716777. doi:10.1080/02724634.2019.1716777. S2CID214520220.
^Oliver A. López-Conde; Adán Pérez-García; María L. Chavarría-Arellano; Jesús Alvarado-Ortega (2020). "A new bothremydid turtle (Pleurodira) from the Olmos Formation (upper Campanian) of Coahuila, Mexico". Cretaceous Research. 119: Article 104710. doi:10.1016/j.cretres.2020.104710. S2CID229448487.
^Ignacio J. Maniel; Marcelo S. de la Fuente; Sebastian Apesteguía; Joaquin Pérez Mayoral; Maria L. Sanchez; Gonzalo D. Veiga; Ian Smales (2020). "Cranial and postcranial remains of a new species of Prochelidella (Testudines: Pleurodira: Chelidae) from 'La Buitrera' (Cenomanian of Patagonia, Argentina), with comments on the monophyly of this extinct chelid genus from southern Gondwana". Journal of Systematic Palaeontology. 18 (12): 1033–1055. Bibcode:2020JSPal..18.1033M. doi:10.1080/14772019.2020.1721579. S2CID214500365.
^Hans Volker Karl, Ulrich Staesche, Amtyaz Safi New findings of neogene tortoises Titanochelon kayadibiensis sp. nov. and Protestudo bessarabica (Riabinin, 1918) (Testudinidae) from the Miocene of western Turkey, with a review of fossil turtles of Turkey SPC Journal of Environmental Sciences, 3 (1) (2021) 1-9
^Juliana Sterli; Ricardo N. Martínez; Ignacio A. Cerda; Cecilia Apaldetti (2020). "Appearances can be deceptive: bizarre shell microanatomy and histology in a new Triassic turtle (Testudinata) from Argentina at the dawn of turtles". Papers in Palaeontology. 7 (2): 1097–1132. doi:10.1002/spp2.1334. S2CID225515560.
^Carolina Oriozabala; Juliana Sterli; Marcelo S. De La Fuente (2020). "New species of the long-necked chelid Yaminuechelys from the Upper Cretaceous (Campanian–Maastrichtian) of Chubut, Argentina". Cretaceous Research. 106: Article 104197. Bibcode:2020CrRes.10604197O. doi:10.1016/j.cretres.2019.104197. S2CID201336536.
^Marcelo S. de la Fuente; Juliana Sterli; Verónica Krapovickas (2020). "Triassic turtles from Pangea: The legacy from South America". Journal of South American Earth Sciences. 105: Article 102910. doi:10.1016/j.jsames.2020.102910. S2CID224891876.
^Tomasz Szczygielski (2020). "Obscure by name: solving the enigma of Chelytherium obscurum, the first described Triassic turtle". Zoological Journal of the Linnean Society. 192 (4): 1111–1122. doi:10.1093/zoolinnean/zlaa139.
^Michael D. Cherney; Jeffrey A. Wilson Mantilla; Iyad Zalmout; Mohammed Sameh M. Antar; Philip D. Gingerich (2020). "New specimens of the Late Eocene turtle Cordichelys (Pleurodira: Podocnemididae) from Wadi Al Hitan and Qasr El-Sagha in the Fayum Province of Egypt". Contributions from the Museum of Paleontology, University of Michigan. 33 (2): 29–64. hdl:2027.42/163364.
^Evangelos Vlachos (2020). "The turtles from 'Gilmore's slab': evolution and diversity of the extinct turtle Echmatemys (Testudines: Testudinoidea: Geoemydidae) from the central United States". Journal of Systematic Palaeontology. 18 (14): 1193–1216. Bibcode:2020JSPal..18.1193V. doi:10.1080/14772019.2020.1754938. S2CID219056552.
^Brenen M. Wynd; Sterling J. Nesbitt; Michelle R. Stocker; Andrew B. Heckert (2020). "A detailed description of Rugarhynchos sixmilensis, gen. et comb. nov. (Archosauriformes, Proterochampsia), and cranial convergence in snout elongation across stem and crown archosaurs". Journal of Vertebrate Paleontology. 39 (6): e1748042. doi:10.1080/02724634.2019.1748042. S2CID219917329.
^M. Jimena Trotteyn; Martín D. Ezcurra (2020). "Redescription of the holotype of Chanaresuchus bonapartei Romer, 1971 (Archosauriformes: Proterochampsidae) from the Upper Triassic rocks of the Chañares Formation of north-western Argentina". Journal of Systematic Palaeontology. 18 (17): 1415–1443. Bibcode:2020JSPal..18.1415T. doi:10.1080/14772019.2020.1768167. S2CID221058509.
^Debajit Datta; Debarati Mukherjee; Sanghamitra Ray (2020). "Taphonomic signatures of a new Upper Triassic phytosaur (Diapsida, Archosauria) bonebed from India: aggregation of a juvenile-dominated paleocommunity". Journal of Vertebrate Paleontology. 39 (6): e1726361. doi:10.1080/02724634.2019.1726361. S2CID219087095.
^Debajit Datta; Kanishak Sharma; Sanghamitra Ray (2020). "Cranial evolution of the Late Triassic phytosaurs (Diapsida, Archosauria): preliminary observations from landmark-based morphometric analysis". Historical Biology: An International Journal of Paleobiology. 33 (11): 2683–2705. doi:10.1080/08912963.2020.1822831. S2CID224916570.
^ abTim Hamley; Juan C. Cisneros; Ross Damiani (2020). "A procolophonid reptile from the Lower Triassic of Australia". Zoological Journal of the Linnean Society. 192 (2): 554–609. doi:10.1093/zoolinnean/zlaa056.
^Paulo R. Romo de Vivar; Agustín G. Martinelli; Annie Schmaltz Hsiou; Marina Bento Soares (2020). "A new rhynchocephalian from the Late Triassic of southern Brazil enhances eusphenodontian diversity". Journal of Systematic Palaeontology. 18 (13): 1103–1126. Bibcode:2020JSPal..18.1103R. doi:10.1080/14772019.2020.1732488. S2CID216226211.
^Hans-Dieter Sues; Rainer R. Schoch (2020). "A new early-diverging sphenodontian (Lepidosauria, Rhynchocephalia) from the Upper Triassic of Virginia, U.S.A.". Journal of Paleontology. 95 (2): 344–350. doi:10.1017/jpa.2020.87. S2CID228808997.
^Zhi-Heng Li; Wei Wang; Han Hu; Ming Wang; HongYu Yi; Jing Lu (2020). "Reanalysis of Oculudentavis shows it is a lizard". Vertebrata PalAsiatica. 59 (2): 95–105. doi:10.19615/j.cnki.1000-3118.201020.
^Fabio Marco Dalla Vecchia (2020). "Raibliania calligarisi gen. n., sp. n., a new tanystropheid (Diapsida, Tanystropheidae) from the Upper Triassic (Carnian) of northeastern Italy". Rivista Italiana di Paleontologia e Stratigrafia. 126 (1): 197–222. doi:10.13130/2039-4942/13041.
^Matthew Skinner; David I. Whiteside; Michael J. Benton (2020). "Late Triassic island dwarfs? Terrestrial tetrapods of the Ruthin fissure (South Wales, UK) including a new genus of procolophonid". Proceedings of the Geologists' Association. 131 (5): 535–561. Bibcode:2020PrGA..131..535S. doi:10.1016/j.pgeola.2020.04.005. S2CID221878712.
^Paulo R. Romo De Vivar; Agustín G. Martinelli; Pedro Henrique M. Fonseca; Marina Bento Soares (2020). "To be or not to be: the hidden side of Cargninia enigmatica and other puzzling remains of Lepidosauromorpha from the Upper Triassic of Brazil". Journal of Vertebrate Paleontology. 40 (4): e1828438. Bibcode:2020JVPal..40E8438R. doi:10.1080/02724634.2020.1828438. S2CID227249876.
^Sofia A. V. Chambi-Trowell; David I. Whiteside; Michael J. Benton; Emily J. Rayfield (2020). "Biomechanical properties of the jaws of two species of Clevosaurus and a reanalysis of rhynchocephalian dentary morphospace". Palaeontology. 63 (6): 919–939. Bibcode:2020Palgy..63..919C. doi:10.1111/pala.12493. S2CID220902843.
^Silvio Renesto; Evelyn Kustatscher; Piero Gianolla (2020). "A putative juvenile specimen of Eusaurosphargis dalsassoi from the Anisian (Middle Triassic) of Piz da Peres (Dolomites, northern Italy)". Rivista Italiana di Paleontologia e Stratigrafia. 126 (2): 249–259. doi:10.13130/2039-4942/13222.
^Martín D. Ezcurra; Felipe C. Montefeltro; Felipe L. Pinheiro; M. Jimena Trotteyn; Adriel R. Gentil; Oscar E.R. Lehmann; Luciano A. Pradelli (2020). "The stem-archosaur evolutionary radiation in South America". Journal of South American Earth Sciences. 105: Article 102935. doi:10.1016/j.jsames.2020.102935. hdl:11336/144048. S2CID225139967.
^Torsten M. Scheyer; Wei Wang; Chun Li; Feiko Miedema; Stephan N. F. Spiekman (2020). "Osteological re-description of Macrocnemus fuyuanensis (Archosauromorpha, Tanystropheidae) from the Middle Triassic of China". Vertebrata PalAsiatica. 58 (3): 169–187. doi:10.19615/j.cnki.1000-3118.200525.
^Nikolay G. Zverkov; Dmitry V. Grigoriev; Igor G. Danilov (2020). "Early Jurassic palaeopolar marine reptiles of Siberia". Geological Magazine. 158 (7): 1305–1322. doi:10.1017/S0016756820001351. S2CID234428429.
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