Cricetulodon MN 5 (biostratigraphic zone) MN 4 (biostratigraphy) María Dolores Soria Mayor Orleanian Somosaguas fossil site Emiliano Aguirre Sansan paleontological site 2016 in paleomammalogy 2010 in paleomammalogy 2014 in paleomammalogy 2019 in paleomammalogy 2017 in paleomammalogy 2020 in paleomammalogy