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Both families contain ellagic acid. Phloem cells contain a large number of plastids and the leaves have dented borders.
The plants are dioecious and possess small flowers without perianth, and the stigma is, at least weakly, secretory. Gunnerale characters shared with the core of the eudicots are cyanogenesis via phenylalanine, metabolic pathways of isoleucine or valine, presence of the DNA sequence of PI-dB motif, 9 and is common to suffer a small deletion in the sequence of 18S ribosomal DNA. The characters which it shares with the core of eudicotyledons, and also with Buxales and Trochodendrales, are an absence of benzylisoquinoline alkaloids, euAP3 + TM6 genes (gene duplication paleoAP3: Class B), and a loss of the mitochondrial gene rps2.
The Gunneraceae are a mesophilic herb (often oversized), and the hydathodes are well developed and secrete mucilage or perhaps a resinous coating.
The Myrothamnaceae are a reviviscent shrub of arid habitats, and the hydathodes are poorly developed and secrete plant resin.
Both have flowers without perianth, but the details of pollen (e.g. Zavada and Dilcher see 1986 10, Wanntorp et al. 2004th 2004b 11 and 12) differ. In Wilkinson 2000 13 is a table of differences.
