Tar Heel/Coachman Formation
The Tar Heel Formation , also known as the Coachman Formation in South Carolina, is a Late Cretaceous (early to middle Campanian -aged) geologic formation in North Carolina and South Carolina , USA. It preserves fossils , including amber dating back to the Cretaceous period . A locality known as Phoebus Landing, has been dated to 78.5-77.1 Ma, and the formation has been overall dated to the early Campanian based on fossil pollen.[ 1] [ 2] [ 3]
Likely deposited in a nearshore coastal environment representing a lower shoreface , it contains a high diversity of vertebrate remains. It has one of the most diverse dinosaur faunas known from the former landmass of Appalachia , the majority of which are known from two sites: Phoebus Landing along the Cape Fear River in Bladen County, North Carolina , as well as Stokes Quarry in Darlington County, South Carolina .[ 4] [ 5] [ 6]
Fossil pollen grains suggest a subtropical to warm, moist temperate climate for the region, with an ecosystem largely dominated by flowering plants .[ 3] The Tar Heel/Coachman Formation appears to be roughly concurrent with the Marshalltown Formation of New Jersey , which preserves a similar fauna.[ 4]
Paleobiota
Cartilaginous fish
Based on the Paleobiology Database & Robb (1989):[ 7] [ 8]
Cartilaginous fish of the Tar Heel/Coachman Formation
Genus
Species
Location
Notes
Images
"Asteracanthus "
A. sp.
Phoebus Landing, NC
A hybodontid shark.
Brachyrhizodus
B. wichitaensis
Phoebus Landing, NC
An eagle ray .
Carcharias
C. holmdelensis
Phoebus Landing, NC
A sand shark .
C. samhammeri
"Hypolophus "
H. sp.
Phoebus Landing, NC
A whiptail stingray .
Meristodonoides
M. montanensis
Phoebus Landing, NC
A hybodontid shark.
Rhombodus
R. laevis
Phoebus Landing, NC
A rhombodontid ray.
Ischyrhiza
I. mira
Phoebus Landing, NC
A sawskate .
Ischyodus
I . cf. bifurcatus
Phoebus Landing, NC
A callorhinchid chimaera .
Scapanorhynchus
S. texanus
Phoebus Landing, NC
A goblin shark .
Squalicorax
S. kaupi
Phoebus Landing, NC
Stokes Quarry, SC
An anacorid shark. Includes a coprolite potentially assignable to S. kaupi , containing the vertebrae of a baby turtle.[ 9]
S. pristodontus
Squatina
S. hassei
Phoebus Landing, NC
An angelshark .
Ray-finned fish
Based on the Paleobiology Database, Robb (1989) & Stringer et al (2018). Some species are known only from otolith remains from Blue Banks & Auger Hole Landings, North Carolina, which are indicated:[ 7] [ 8] [ 10]
Genus
Species
Location
Material
Notes
Images
Albula
?A. campaniana
Blue Banks & Auger Hole Landings, NC
263-378 otoliths
A bonefish .
?A. cf. A. ripleyensis
Blue Banks Landing, NC
1 otolith
A. sp.
Phoebus Landing, NC
Anomoeodus
A. phaseolus
Phoebus Landing, NC
A pycnodontid pycnodont .
Ariidae indet.
Blue Banks Landing, NC
11 otoliths
A sea catfish of uncertain affinities.
Aulopidae indet.
Blue Banks Landing, NC
1 otolith
A flagfin of uncertain affinities.
Beryx
?B. maastrichtiensis
Blue Banks & Auger Hole Landings, NC
55 otoliths
A potential relative of alfonsinos .
?B. zideki
41 otoliths
?Congridae indet.
Auger Hole Landing, NC
1 otolith
An apparent conger eel of uncertain affinities.
Cylindracanthus
C. ornatus
Phoebus Landing, NC
A fish of uncertain affinities.
Enchodus
E. cf. petrosus
Phoebus Landing, NC
An enchodontid aulopiform .
Gonostomatidae indet.
Blue Banks & Auger Hole Landings, NC
4 otoliths
A bristlemouth of uncertain affinities.
Hoplostethus
?H. coffeesandensis
Blue Banks & Auger Hole Landings, NC
4 otoliths
A slimehead .
Kokenichthys
K. ensis
Blue Banks Landing, NC
2 otoliths
An elopiform or osteoglossiform .[ 11]
Lepisosteidae indet.
Phoebus Landing, NC
A gar .
Megalopidae indet.
Blue Banks Landing, NC
1 otolith
A tarpon of uncertain affinities.
Osmeroides
O. weileri
Blue Banks Landing, NC
6 otoliths
An osmeroidid elopomorph .
Paralbula
P. casei
Phoebus Landing, NC
A phyllodontid elopomorph.
?Pempheris
?P. huddlestoni
Auger Hole Landing, NC
1 otolith
An apparent sweeper .
Percoidei indet.
Blue Banks & Auger Hole Landings, NC
5 otoliths
A percoid .
?Polymixia
?P. cf. harderi
Blue Banks Landing, NC
An apparent beardfish .
Pterothrissus
P. carolinensis
Blue Banks Landing, NC
67 otoliths
A pterothrissine bonefish.
Saurodon
S. sp.
Phoebus Landing, NC
A saurodontid ichthyodectiform.
Stephanodus
S. sp.
Phoebus Landing, NC
A pycnodont.
Xiphactinus
X. audax
Phoebus Landing, North Carolina, NC
An ichthyodectid ichthyodectiform.
X. vetus
Reptiles
Records from SC based on Schwimmer et al (2015):[ 6]
Dinosaurs
Ornithischians
Based mainly on Brownstein (2018):[ 4]
Ornithischians of the Tar Heel/Coachman Formation
Genus
Species
Location
Notes
Images
cf. Hadrosaurus
?H. foulkii
Phoebus Landing & Sampson County, NC[ 12]
A hadrosaurid , tentatively assigned to the well-known H. foulkii .
"Hadrosaurus "
"H." minor
Phoebus Landing, NC
A small or juvenile hadrosaur, nomen dubium .
Hypsibema
H. crassicauda
Phoebus Landing & Sampson County, NC
A gigantic hadrosauroid . First discovered in North Carolina from Sampson County , 1869, and described by Edward Drinker Cope . It was measured to be 12-17 meters, making it one of the largest hadrosauroids.[ 13] Type locality for genus and species.
Leptoceratopsidae indet.
Sampson County, NC
A leptoceratopsid ceratopsian . Known from an indeterminate left maxilla found in 2016.[ 14]
Lophorhothon
L. atopus
Phoebus Landing, NC
A hadrosauromorph . Originally described from the Mooreville Chalk Formation , Alabama .
An indeterminate hadrosauroid is known from Stokes Quarry, SC.[ 4]
Theropods
Indeterminate theropods, ornithomimosaurs, and maniraptorans are known from Stokes Quarry.[ 4]
Crocodylomorphs
Based on the Paleobiology Database:[ 7]
Genus
Species
Location
Notes
Images
Borealosuchus
B. formidabilis
Phoebus Landing, NC
A eusuchian .
Bottosaurus
B. sp.
Stokes Quarry, SC
An early caiman .
Deinosuchus
D. rugosus (=Thecachampsa rugosa , Polydectes biturgidus , Polyptychodon rugosus )
Phoebus Landing & Clifton Farm, NC
Stokes Quarry, SC
An alligatoroid , one of the largest known crocodilians. Type locality for genus and species.
Gavialoidea indet.
Stokes Quarry, SC
A gavialoid .
Turtles
Based on the Paleobiology Database:[ 7]
Genus
Species
Location
Notes
Images
Adocus
A. beatus
Phoebus Landing, NC
Stokes Quarry, SC
An adocid .
Bothremys
B. cooki
Phoebus Landing, NC
A bothremydid side-necked turtle .
Chedighaii
C. barberi
Phoebus Landing, NC
A bothremydid side-necked turtle.
C. hutchinsoni
Corsochelys
C. bentlyi
Stokes Quarry, SC
A sea turtle , likely a dermochelyid . Type locality for species.
Euclastes
E. wielandi
Stokes Quarry, SC
A pancheloniid sea turtle.
Osteopygis
O. emarginatus
Phoebus Landing, NC
A macrobaenid .
?Taphrosphys
?T. dares
Phoebus Landing, NC
A bothremydid side-necked turtle, taxonomic placement disputed.
Toxochelys
T. sp.
Stokes Quarry, SC
A toxochelyid sea turtle.
"Trionyx "
"T ." halophilus
Phoebus Landing, NC
A stem -softshell turtle , likely not an actual member of the genus Trionyx . Species names are nomina dubia .
"T. " priscus
Stokes Quarry, SC
Plesiosaurs
Squamates
Mammals
Based on the Paleobiology Database:[ 20]
References
^ Longrich, Nicholas R. (2016). "A ceratopsian dinosaur from the Late Cretaceous of eastern North America, and implications for dinosaur biogeography" . Cretaceous Research . 57 : 199– 207. Bibcode :2016CrRes..57..199L . doi :10.1016/j.cretres.2015.08.004 .
^ "Geolex — TarHeel publications" . ngmdb.usgs.gov . Retrieved 2024-11-13 .
^ a b Mitra, Madhumi (2002-07-29). Paleopalynology of the Tar Heel Formation of Atlantic Coastal Plain of North Carolina, United States (Thesis).
^ a b c d e f g h i Brownstein, Chase D. (2018-02-08). "The biogeography and ecology of the Cretaceous non-avian dinosaurs of Appalachia" . Palaeontologia Electronica . 21 (1): 1– 56. doi :10.26879/801 . ISSN 1094-8074 .
^ "PBDB Collection" . paleobiodb.org . Retrieved 2024-11-13 .
^ a b Schwimmer, David R.; Sanders, Albert E.; Erickson, Bruce R.; Weems, Robert E. (2015). "A Late Cretaceous Dinosaur and Reptile Assemblage from South Carolina, Usa" . Transactions of the American Philosophical Society . 105 (2): i–157. ISSN 0065-9746 .
^ a b c d Czaplewski, John J. "PBDB Navigator" . paleobiodb.org . Retrieved 2024-10-10 .
^ a b Albert J Robb III (1989). The Upper Cretaceous (Campanian, Black Creek Formation) fossil fish fauna of Phoebus Landing, Bladen County, North Carolina .
^ Schwimmer, David R.; Weems, Robert E.; Sanders, Albert E. (2015). "A Late Cretaceous Shark Coprolite with Baby Freshwater Turtle Vertebrae Inclusions" . PALAIOS . 30 (9/10): 707– 713. ISSN 0883-1351 .
^ Stringer, Gary L.; Clements, Don; Sadorf, Eric; Shannon, Kevin (2019). "First Description and Significance of Cretaceous Teleostean Otoliths (Tar Heel Formation, Campanian) from North Carolina" . Eastern Paleontologist . 4 : 1– 22. ISSN 2475-5117 .
^ Stringer, Gary L.; Sloan, James Carson (2023-05-25). "First Cretaceous teleostean otolith assemblage (Arkadelphia Formation, upper Maastrichtian) from Arkansas, USA, early Gadiformes, and the Western Interior Seaway" . PaleoBios . 40 (3). doi :10.5070/P940361192 . ISSN 2373-8189 .
^ NCSM Paleontology Database
^ Miller, Halsey W. (1967). "Cretaceous Vertebrates from Phoebus Landing, North Carolina" . Proceedings of the Academy of Natural Sciences of Philadelphia . 119 : 219– 239. ISSN 0097-3157 .
^ Longrich, Nicholas R. (2016-01-01). "A ceratopsian dinosaur from the Late Cretaceous of eastern North America, and implications for dinosaur biogeography" . Cretaceous Research . 57 : 199– 207. doi :10.1016/j.cretres.2015.08.004 . ISSN 0195-6671 .
^ Baird D., and Horner, J., 1979, "Cretaceous dinosaurs of North Carolina" Brimleyana 2 : 1–28
^ Brownstein, Chase D. (2018-12-01). "A large dromaeosaurid from North Carolina" . Cretaceous Research . 92 : 1– 7. Bibcode :2018CrRes..92....1B . doi :10.1016/j.cretres.2018.07.006 . ISSN 0195-6671 . S2CID 135459468 .
^ Mortimer, Mickey (2024). "Tyrannosauroidea" . The Theropod Database . Retrieved Jan 28, 2025 . {{cite web }}
: CS1 maint: url-status (link )
^ Brownstein, Chase D. (2018-04-05). "Diversity of raptor dinosaurs in southeastern North America revealed by the first definite record from North Carolina" . PeerJ Preprints . doi :10.7287/peerj.preprints.26829v1 .
^ Brownstein, Chase D. (2018). "The distinctive theropod assemblage of the Ellisdale site of New Jersey and its implications for North American dinosaur ecology and evolution during the Cretaceous" . Journal of Paleontology . 92 (6): 1115– 1129. Bibcode :2018JPal...92.1115B . doi :10.1017/jpa.2018.42 . ISSN 0022-3360 .
^ a b c "PBDB Collection" . paleobiodb.org . Retrieved 2024-11-15 .