Helice tridens

Helice tridens
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Malacostraca
Order:	Decapoda
Suborder:	Pleocyemata
Infraorder:	Brachyura
Family:	Varunidae
Genus:	Helice
Species:	H. tridens
Binomial name
	Helice tridens; (De Haan, 1835)
Synonyms
	Ocypode tridens De Haan, 1835; Cyclograpsus latreillii H. Milne-Edwards, 1837; Helice latreillei H. Milne-Edwards, 1837;

Helice tridens is a species of crab which lives on mudflats around the coasts of Japan and the Korean Peninsula.^[3]

Ecology

It is semi-terrestrial, returning to the sea to spawn.^[4] The species appears to be adversely affected by the presence of raccoons (Procyon lotor), an invasive predator.^[4] H. tridens has a salinity requirement which lies between those of two other estuarine crabs in Japan, Helicana japonica and Chiromantes dehaani.^[5]

Smaller individuals shelter in burrows in reed marshes, apparently in order to avoid cannibalism; this may also be the reason for the migration of larger individuals to brackish water lagoons in summer, when the crabs exceed their carrying capacity.^[6]

Taxonomy

Helice tridens was first described by Wilhem de Haan in an 1835 volume of Fauna Japonica, as Ocypode tridens.^[7] The former subspecies H. t. wuana and H. t. sheni are now recognised as a separate species, Helicana wuana.^[2]

References

^ "Helice tridens". Crabs of Japan. Retrieved January 31, 2009.
^ ^a ^b P. K. L. Ng, D. Guinot & P. J. F. Davie (2008). "Systema Brachyurorum: Part I. An annotated checklist of extant Brachyuran crabs of the world" (PDF). Raffles Bulletin of Zoology. 17: 1–286. Archived from the original (PDF) on 2011-06-06.
^ Hsi-Te Shih & Hiroshi Suzuki (2008). "Taxonomy, phylogeny, and biogeography of the endemic mudflat crab Helice/Chasmagnathus complex (Crustacea: Brachyura: Varunidae) from East Asia" (PDF). Zoological Studies. 47 (1): 114–125. Archived from the original (PDF) on 2017-08-09. Retrieved 2010-03-27.
^ ^a ^b Hisayo Hayama, Masato Kaneda & Mayuh Tabata (2006). "Rapid range expansion of the feral raccoon (Procyon lotor) in Kanagawa Prefecture, Japan, and its impact on native organisms". In F. Koike; M. N. Clout; M. Kawamichi; M. De Poorter & K. Iwatsuki (eds.). Assessment and Control of Biological Invasion Risks (PDF). Shoukadoh Book Sellers, Kyoto, Japan and IUCN, Gland, Switzerland. pp. 196–199. Archived from the original (PDF) on 2011-07-22. Retrieved 2009-01-31.
^ Bambang Irawan & Akihiro Kijima (1994). "Difference of salinity requirements among the three estuarine crab species, Chiromantes dehaani, Helice tridens and H. japonica (Brachyura: Grapsidae)" (PDF). Tohoku Journal of Agricultural Research. 44 (1–4). Archived from the original (PDF) on 2012-06-02. Retrieved 2009-01-31.
^ Yasushi Kurihara, Katsuhiro Sekimoto & Masahide Miyata (1988). "Wandering behaviour of the mud-crab Helice tridens related to evasion of cannibalism" (PDF). Marine Ecology Progress Series. 49: 41–50. Bibcode:1988MEPS...49...41K. doi:10.3354/meps049041.
^ Wilhem de Haan (1835). "Ocypode (Helice) tridens sp. nov.". In Philipp Franz von Siebold (ed.). Fauna Japonica. p. 57. Archived from the original on 2011-07-25. Retrieved 2009-01-31.

External links

Media related to Helice tridens at Wikimedia Commons

[1] "Helice tridens". Crabs of Japan. Retrieved January 31, 2009.

[Ng-2] P. K. L. Ng, D. Guinot & P. J. F. Davie (2008). "Systema Brachyurorum: Part I. An annotated checklist of extant Brachyuran crabs of the world" (PDF). Raffles Bulletin of Zoology. 17: 1–286. Archived from the original (PDF) on 2011-06-06.

[3] Hsi-Te Shih & Hiroshi Suzuki (2008). "Taxonomy, phylogeny, and biogeography of the endemic mudflat crab Helice/Chasmagnathus complex (Crustacea: Brachyura: Varunidae) from East Asia" (PDF). Zoological Studies. 47 (1): 114–125. Archived from the original (PDF) on 2017-08-09. Retrieved 2010-03-27.

[Hayama-4] Hisayo Hayama, Masato Kaneda & Mayuh Tabata (2006). "Rapid range expansion of the feral raccoon (Procyon lotor) in Kanagawa Prefecture, Japan, and its impact on native organisms". In F. Koike; M. N. Clout; M. Kawamichi; M. De Poorter & K. Iwatsuki (eds.). Assessment and Control of Biological Invasion Risks (PDF). Shoukadoh Book Sellers, Kyoto, Japan and IUCN, Gland, Switzerland. pp. 196–199. Archived from the original (PDF) on 2011-07-22. Retrieved 2009-01-31.

[5] Bambang Irawan & Akihiro Kijima (1994). "Difference of salinity requirements among the three estuarine crab species, Chiromantes dehaani, Helice tridens and H. japonica (Brachyura: Grapsidae)" (PDF). Tohoku Journal of Agricultural Research. 44 (1–4). Archived from the original (PDF) on 2012-06-02. Retrieved 2009-01-31.

[6] Yasushi Kurihara, Katsuhiro Sekimoto & Masahide Miyata (1988). "Wandering behaviour of the mud-crab Helice tridens related to evasion of cannibalism" (PDF). Marine Ecology Progress Series. 49: 41–50. Bibcode:1988MEPS...49...41K. doi:10.3354/meps049041.

[7] Wilhem de Haan (1835). "Ocypode (Helice) tridens sp. nov.". In Philipp Franz von Siebold (ed.). Fauna Japonica. p. 57. Archived from the original on 2011-07-25. Retrieved 2009-01-31.

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Helice tridens

Ecology

Taxonomy

References

External links

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