Maip

Maip; Temporal range: Late Cretaceous, Maastrichtian PreꞒ Ꞓ O S D C P T J K Pg N
	Known material (A), reconstruction of the thoracic cavity at level of sixth dorsal vertebra (B), and interpretative drawing of the excavation (C)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	Saurischia
Clade:	Theropoda
Clade:	†Megaraptora
Family:	†Megaraptoridae
Genus:	†Maip; Rolando et al., 2022
Species:	†M. macrothorax
Binomial name
	†Maip macrothorax; Rolando et al., 2022

Maip is a genus of large megaraptorid theropod dinosaur from the Late Cretaceous (Maastrichtian) Chorrillo Formation of Santa Cruz, Argentina. The genus contains a single species, M. macrothorax, known from an incomplete, disarticulated skeleton. Maip may represent the largest megaraptorid known from South America, and possibly the world.^[1]

Discovery and naming

The Maip holotype specimen, MPM 21545, was discovered by Alexis Rolando on the La Anita Farm, 30 kilometres (19 mi) west of El Calafate, Santa Cruz province, Argentina, in 2019. The specimen was found disarticulated but in association over an area of 5 by 3 square metres (54 sq ft × 32 sq ft). The known fossil material consists of the axis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsal. Some of these bones were described in 2019 by Novas et al.^[2] The holotype specimen includes some bones previously unknown in other megaraptorids. Although fragmentary, it represents one of the most complete megaraptorid skeletons known.^[1]

Maip was first announced in a Research Square preprint in 2021. However, since the paper did not meet the necessary requirements, the taxon was considered informally named.^[3] In 2022, the fossil material was validly described as belonging to a new genus and species of megaraptorid by Rolando et al. The generic name, "Maip", references a malicious being in Aonikenk mythology that is "the shadow of death" that "kills with cold wind." The specific name, "macrothorax", is derived from the Greek "makrós", meaning "large", and the Latin "thorax", meaning "chest", in reference to its large thoracic cavity.^[1]

Description

Maip is estimated to have been around 9–10 metres (30–33 ft) long in life. As such, it may be the largest megaraptorid currently known. Members of the Megaraptoridae as a whole increased in body length following the extinction of the carcharodontosaurids in the Southern Hemisphere in the early Late Cretaceous.^[4] It is hypothesized that the absence of these large apex predators allowed other theropods to diversify and fill the empty niche space. Megaraptorids, in addition to abelisaurids and unenlagiids, then became the primary predators in their ecosystems. An analysis of known megaraptorans shows that members of the clade in Asia, South America, and Australia ranged from 4–4.5 metres (13–15 ft) long during the Barremian-Aptian, after which Aptian-early Turonian forms increased to 4.5–6 metres (15–20 ft) in Australia and South America. Turonian-Coniacian megaraptorids, only known from South America, were larger, at 6–7 metres (20–23 ft) long, and the final Santonian-Maastrichtian forms were larger still, at 7–10 metres (23–33 ft).^[1]

Traces on the preserved ribs were interpreted by the describing authors as being attachment points for ligaments. From these, it was inferred that Maip would have likely had a respiratory system comparable to those of modern birds, rather than those of other extant reptiles, such as crocodilians.^[1]

Classification

In their phylogenetic analyses, Rolando et al. (2022) recovered Maip as a derived megaraptorid in a polytomy with other Argentinian megaraptorids. They also noted the presence of two distinct clades: a more inclusive clade, comprising all megaraptorids except Fukuiraptor and Australovenator, (shown below as "Clade A"), and a more exclusive clade of larger, entirely South American megaraptorids (shown below as "Clade B"). Like some previous analyses by other authors, Megaraptora is nested within Coelurosauria, as the sister taxon to Tyrannosauroidea. The cladogram below displays the megaraptoran results of the phylogenetic analyses by Rolando et al (2022).^[1]

Australian megaraptorid indet. (LRF 100–106)

Australovenator

"Clade A"

Aoniraptor

Bajo Barreal Formation megaraptorid indet. (UNPSJB-Pv 944/958)

Megaraptor

Murusraptor

"Clade B"

	Orkoraptor

	Tratayenia

	Maip

	Aerosteon

Paleoecology

Maip is known from the Maastrichtian-dated Chorrillo Formation of southern Argentina. Other named dinosaurs known from the formation include Isasicursor, an elasmarian ornithopod, and Nullotitan, a titanosaurian sauropod.^[2] Indeterminate remains belonging to ankylosaurs, euiguanodontians, hadrosaurids, noasaurids, and unenlagiids have also been recovered from the formation. Fossils of indeterminate anurans, fish, mammals, mosasaurs, snakes, turtles, and gastropods are also known.^[5] Very fragmentary fossil material, including teeth and a dorsal centrum, has been classified as having megaraptorid affinities, but it is too fragmentary to be assigned to Maip or any other taxon.^[1]

References

^ ^a ^b ^c ^d ^e ^f ^g Rolando, Alexis M. A.; Motta, Matias J.; Agnolín, Federico L.; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (April 26, 2022). "A large Megaraptoridae (Theropoda: Coelurosauria) from Upper Cretaceous (Maastrichtian) of Patagonia, Argentina". Scientific Reports. 12 (1): Article number 6318. Bibcode:2022NatSR..12.6318A. doi:10.1038/s41598-022-09272-z. PMC 9042913. PMID 35474310.
^ ^a ^b Novas, Fernando; Agnolin, Federico; Rozadilla, Sebastián; Aranciaga-Rolando, Alexis; Brissón-Eli, Federico; Motta, Matias; Cerroni, Mauricio; Ezcurra, Martín; Martinelli, Agustin; D´Angelo, Julia; Álvarez-Herrera, Gerardo (2019). "Paleontological discoveries in the Chorrillo Formation (upper Campanian-lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentina". Revista del Museo Argentino de Ciencias Naturales. Nueva Serie. 21 (2): 217–293. doi:10.22179/revmacn.21.655. hdl:11336/122229. ISSN 1853-0400. S2CID 212808625.
^ Rolando, Alexis M. A.; Motta, Matias J.; Agnolín, Federico L.; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (December 22, 2021). "The Biggest Megaraptoridae (Theropoda: Coelurosauria) of South America" (PDF). Research Square: 1–43. doi:10.21203/rs.3.rs-1152394/v1.
^ Meso, J. G.; Juárez Valieri, R. D.; Porfiri, J. D.; Correa, S. A. S.; Martinelli, A. G.; Casal, G. A.; Canudo, J. I.; Poblete, F.; Dos Santos, D. (September 1, 2021). "Testing the persistence of Carcharodontosauridae (Theropoda) in the Upper Cretaceous of Patagonia based on dental evidence". Cretaceous Research. 125: 104875. Bibcode:2021CrRes.12504875M. doi:10.1016/j.cretres.2021.104875. hdl:11336/183886. ISSN 0195-6671.
^ Rozadilla, Sebastián; Agnolín, Federico; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (September 1, 2021). "Ornithischian remains from the Chorrillo Formation (Upper Cretaceous), southern Patagonia, Argentina, and their implications on ornithischian paleobiogeography in the Southern Hemisphere". Cretaceous Research. 125: 104881. Bibcode:2021CrRes.12504881R. doi:10.1016/j.cretres.2021.104881. ISSN 0195-6671.

[:0-1] ^ ^a ^b ^c ^d ^e ^f ^g Rolando, Alexis M. A.; Motta, Matias J.; Agnolín, Federico L.; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (April 26, 2022). "A large Megaraptoridae (Theropoda: Coelurosauria) from Upper Cretaceous (Maastrichtian) of Patagonia, Argentina". Scientific Reports. 12 (1): Article number 6318. Bibcode:2022NatSR..12.6318A. doi:10.1038/s41598-022-09272-z. PMC 9042913. PMID 35474310.

[:1-2] Novas, Fernando; Agnolin, Federico; Rozadilla, Sebastián; Aranciaga-Rolando, Alexis; Brissón-Eli, Federico; Motta, Matias; Cerroni, Mauricio; Ezcurra, Martín; Martinelli, Agustin; D´Angelo, Julia; Álvarez-Herrera, Gerardo (2019). "Paleontological discoveries in the Chorrillo Formation (upper Campanian-lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentina". Revista del Museo Argentino de Ciencias Naturales. Nueva Serie. 21 (2): 217–293. doi:10.22179/revmacn.21.655. hdl:11336/122229. ISSN 1853-0400. S2CID 212808625.

[3] Rolando, Alexis M. A.; Motta, Matias J.; Agnolín, Federico L.; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (December 22, 2021). "The Biggest Megaraptoridae (Theropoda: Coelurosauria) of South America" (PDF). Research Square: 1–43. doi:10.21203/rs.3.rs-1152394/v1.

[4] Meso, J. G.; Juárez Valieri, R. D.; Porfiri, J. D.; Correa, S. A. S.; Martinelli, A. G.; Casal, G. A.; Canudo, J. I.; Poblete, F.; Dos Santos, D. (September 1, 2021). "Testing the persistence of Carcharodontosauridae (Theropoda) in the Upper Cretaceous of Patagonia based on dental evidence". Cretaceous Research. 125: 104875. Bibcode:2021CrRes.12504875M. doi:10.1016/j.cretres.2021.104875. hdl:11336/183886. ISSN 0195-6671.

[5] Rozadilla, Sebastián; Agnolín, Federico; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (September 1, 2021). "Ornithischian remains from the Chorrillo Formation (Upper Cretaceous), southern Patagonia, Argentina, and their implications on ornithischian paleobiogeography in the Southern Hemisphere". Cretaceous Research. 125: 104881. Bibcode:2021CrRes.12504881R. doi:10.1016/j.cretres.2021.104881. ISSN 0195-6671.

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Maip

Discovery and naming

Description

Classification

Paleoecology

References

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