This is an overview of the paleofauna of the EoceneMessel Formation as explored by the Messel Pit excavations in Germany. A former quarry and now UNESCO World Heritage Site, the Messel Formation preserves what once were a series of anoxic lakes surrounded by a sub-tropical rainforest during the Middle Eocene, approximately 47 Ma.
A possibly rheophile species sponge, it grew dominant under favorable conditions and gradually replaced the native population of Lutetiospongilla until a change to the environment. Decrease in freshwater supply likely diminished the sponges population.
A permanent resident of the Messel lake, L. heili was already present when Ephydatia arrived in the lake and differed from the other species by possibly being rheophobe. Following the later species decline, Lutetiospongilla became the second "mass species" of the lake.
An unnamed sponge, informally referred to as "Type 3" by Heil (1964). It may have colonized Messel lake after Lutetiospongilla had replaced Ephydatia as the dominant sponge species.[1]
Thaumaturus is only known from juvenile specimens, ranging in length from 3–6 cm (1.2–2.4 in).[2] Its relationship with other groups of fish remains unknown.
Messelobatrachus and M. tobieni were first coined in a 1988 PhD. dissertation, but never formally published.[43] One fossil preserves a possible tadpole of "Messelobatrachus".
Phylogenetic analysis indicates that it may be a close relative of the limbless Amphisbaenia. Only two specimens are known, one of which was found in the stomach of the varanoid Paranecrosaurus
First described as "Saniwa" feisti, it was found with the skull of Cryptolacerta in its guts. This indicates that this varanoid was at least partly carnivorous.
A small species of alligatorid, some consider it a species of Allognathosuchus. Christopher Brochu argues for the continued use of Hassiacosuchus however.[76]
Its size was intermediate between Lithornis plebius and Lithornis hookeri. Additionally, it represents the last known lithornithid in the fossil record.
A wading bird thought to be basal to both flamingos and grebes. Both the beak shape and the presence of gastroliths show that it was not yet adapted to filderfeed and instead fed on hard items.
An early genus of swift, its reduced legs and less developed wings indicate that it wasn't as aerial as its modern relatives and likely nested in trees.
The most common fossil bird found at Messel, it was likely a terrestrial animal capable of only short bursts of flight due to its short wings. It may have had a comb or crest atop its head.
Originally described as an owl from the Geiseltal, it was later found to be related to mousebirds. Its tail is notably shorter than that of its extant relatives. It is possible that the indetermined species instead belongs to Anneavis.
Relatives of the modern hoopoe and wood hoopoe, they were birds with long, pointed beaks and diminutive body size. The tail feathers of one specimen show they were barred, preserving a pattern of dark and light stripes.
Several groups of Messel birds share characteristics with the modern seriemas, which has led to them being placed within the clade Cariamae in the past. However, this placement typically occurred under the assumption that they are a group within gruiformes, which has been disputed by more recent analysis. Instead more recent publications consider Cariamae (or Cariamiformes) as basal members of Australaves.[112]
Salmila was originally described as a part of Cariamae within Gruiformes, but later analysis suggested that it was a sister taxon to the group instead. It shares characteristics with both seriemas and trumpeter birds.
Messelastur is a bird with a poorly understood ecology. Although shown to be closely related to Tynsyka, its relationship to other groups is not fully understood. Older publications suggest it may have been related to parrots or various birds of prey (including owls, hawks and falcons). One recent publication positions them at the base of Psittacopasserae alongside the Pseudasturidae (=Halcyornithidae).[119]
Zygodactyl birds originally thought to be relatives of woodpeckers, later research placed them as a sister group to the Passeriformes. Diet varries between species, with the type species likely being an insectivore while the larger Primozygodactylus mayor was found with grapes in its stomach. The wings were short and the legs long, indicating a maneuverable animal foraging in the undergrowth. At least in some species the central two tail feathers were greatly elongated and possibly used in display.
A poorly preserved pseudasturid of smaller size than Serudaptus. It may be a different species of the same genus, a juvenile, a member of the opposite sex or a unique genus.
A bird with zygodactyl feet and long claws suited for climbing. Its exact phylogenetic position is poorly understood, with some research indicating it may be a basal member of the clade containing Passeriformes and Psittaciformes.
The genus is named for the bony tubercles present on the cervical vertebrae. Although its relationship is not fully understood, it was found to not be a member of Idiornithidae and shared traits with Anhimidae and Cathartidae.
A basal genus among the Messel bats, Archaeonycteris resembles modern vesper bats in wing morphology and likely preferred more open spaces in the upper trunk areas to hunt. Stomach contents indicate a preference for beetles.[2]
Hassianycteris was a somewhat heavy bat with narrow wings. Based on its anatomy, which resembles modern free-tailed bats, it was a fast flying animal hunting above the treetops.[2]
The wing morphology of Palaeochiropteryx indicates that it flew slowly but agile relatively close to the ground to catch insects such as moths and butterflies.[2]
Buxolestes is a member of a basal group of mammals. The proportions of Buxolestes piscator are similar to modern otters and its stomach contents indicate a diet of fish and small vertebrates. B. minor shows adaptations towards digging and was found with plant material in its stomach, but likely also fed on various animals.
Macrocranion were small, likely nocturnal, animals living on the forest floor. M. tupaiodon reached a size of up to 30 cm (12 in) and was possibly omnivorous with woolly fur.[2] The smaller M. tenerum meanwhile was inferred to be insectivorous and covered in bristles. Some researchers suggest it may have moved like modern springhares.[2][138]
A larger relative of Macrocranion, Pholidocercus was an animal 40 cm (16 in). Its proportions indicate that it was a quadrupedal animal that lived on the forest floor. The tail was covered in overlapping scales and the head was similarly covered in a hardened structure while the torso was covered in thick bristles.[2]
Known from Messel in the form of three species and multiple specimens, Leptictidium was a eutherian mammal of uncertain relationships. Its hindlimbs are noticeably larger than its forelimbs, but its means of locomotion are not agreed upon.
Agile predators of the forest floor, at least one species (L. behnkeae) is known to have had a bushy tail. The exact position hyaenodontids like Lesmesodon have within mammals is debated.
Eomanis is a more derived member of Pholidota than the contemporary genera found in Messel and already shows the classic scales this group is known for.[149]
Originally species of Propalaeotherium and Lophiotherium, they were eleveted to a distinct genus in 2006. Several specimens were preserved with their fetus, showing that they only carried a single foal.
An early primate known from a single specimen preserved in two slabs. The counterslab includes several elements fabricated to make it appear more complete. The classification of Darwinius has been subject to continued debate.
An early primate primarily known from partial remains. One lower jaw was found in the coprolite of Buxolestes while another fossil was found with the tooth of a crocodile embedded into it.[2]
An arboreal rodent that resembles modern squirrels in proportion. It reached a total length of 1 m (3 ft 3 in), most of which consisting of the bushy tail, and primarily fed on leaves.[2]
M. parvus was sometimes placed in Hartenbergeromys,[2] However a 2019 study found the asserted differences to the type species insufficient to warrant this split. Its tail was only sparsely covered in fur.[163]
References
^ abcdRichter, G.; Baszio, S. (2009). "Geographic and stratigraphic distribution of spongillids (Porifera) and the leit value of spiculites in the Messel Pit Fossil Site". Palaeobio Palaeoenv. 89 (1–2): 53–66. doi:10.1007/s12549-009-0003-7. S2CID129838426.
^ abcRichter, G.; Wedmann, S. (2005). "Ecology of the Eocene Lake Messel revealed by analysis of small fish coprolites and sediments from a drilling core". Palaeogeography, Palaeoclimatology, Palaeoecology. 223 (1–2): 147–161. Bibcode:2005PPP...223..147R. doi:10.1016/j.palaeo.2005.04.002.
^Wunderlich, J. (1986). "Die ersten Spinnen aus dem Mittel-Eozän der Grube Messel. Fossilfundstelle Messel, Nr. 49". Senckenbergiana Lethaea. 67 (1–4): 171–176.
^ abSmith, K.T.; Schaal, S.F.K.; Habersetzer, J. (2018). "7 Jewels in the Oil Shale – Insects and Other Invertebrates". Messel-An Ancient Greenhouse Ecosystem. Senckenberg. pp. 67–69.
^Tröster, G. (1999). "An unusual new fossil click-beetle (Coleoptera: Elateridae) from the Middle Eocene of the Grube Messel (Germany)". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. 1999: 11–20. doi:10.1127/njgpm/1999/1999/11.
^ abcdefTröster, G. (1994). "Neue Arten der Gattung Macropunctum (Insecta, Coleoptera, Elateridae) aus der Olschieferfazies der mitteleozänen Messelformation der Grube Messel bei Darmstadt". PalZ. 68 (1–2): 145–162. doi:10.1007/bf02989437. S2CID127993355.
^Šmídová, L.; Vidlička, L.; Wedmann, S. (2021). "Appearance of the family Blaberidae (Insecta: Blattaria) during the Cretaceous and a review of fossils of this family". Palaeontographica Abteilung A. 321 (1–6): 71–79. doi:10.1127/pala/2021/0109. S2CID243467634.
^ abcTorsten Wappler; Ernst Heiss; Sonja Wedmann (2015). "New flatbug (Hemiptera: Heteroptera: Aradidae) records from the Middle Eocene Messel Maar, Germany". Paläontologische Zeitschrift. 89 (3): 653–660. Bibcode:2015PalZ...89..653W. doi:10.1007/s12542-014-0237-1. S2CID83668045.
^ abcSzwedo, J.; Wappler, T. (2006). "New planthoppers (Insecta: Hemiptera: Fulgoromorpha) from the Middle Eocene Messel Maar". Annales Zoologici. 56 (3): 555–566. doi:10.3161/000345406778811907 (inactive 2024-09-19).{{cite journal}}: CS1 maint: DOI inactive as of September 2024 (link)
^ abcWappler, T. (2003). "New fossil lace bugs (Heteroptera: Tingidae) from the Middle Eocene of the Grube Messel (Germany), with a catalog of fossil lace bugs". Zootaxa. 374: 1–26. doi:10.11646/zootaxa.374.1.1.
^Dlussky, GM; Wappler, T; Wedmann, S (2009). "Fossil ants of the genus Gesomyrmex Mayr (Hymenoptera, Formicidae) from the Eocene of Europe and remarks on the evolution of arboreal ant communities". Zootaxa. 2031: 1–20. doi:10.11646/zootaxa.2031.1.1. S2CID56253714.
^Wedmann, S.; Pouillon, J.; Nel, A. (2014). "New Palaeogene horntail wasps (Hymenoptera, Siricidae) and a discussion of their fossil record". Zootaxa. 3869 (1): 33–43. doi:10.11646/zootaxa.3869.1.3. PMID25283896.
^Abels, J.; Wedmann, S. (2021). "A fossil wasp (Hymenoptera: Vespidae: Vespula? hassiaca) from the Eocene of Messel". Palaeontographica Abteilung A. 323 (1–3): 105–117. doi:10.1127/pala/2021/0122. S2CID245248645.
^ abRomain Garrouste; André Nel (2015). "New Eocene damselflies and first Cenozoic damsel-dragonfly of the isophlebiopteran lineage (Insecta: Odonata)". Zootaxa. 4028 (3): 354–366. doi:10.11646/zootaxa.4028.3.2. PMID26624314.
^Grande, Lance (2010). "An empirical synthetic pattern study of gars (Lepisosteiformes) and closely related species, based mostly on skeletal anatomy. The resurrection of Holostei". Ichthyology & Herpetology. 10.
^Micklich, N.; Klappert, G. (2004). "Character variation in some Messel fishes". Mesozoic Fishes 3: 137–163.
^Micklich, N. (1978). "Palaeoperca proxima, ein neuer Knochenfisch aus dem Mittel-Eozän von Messel bei Darmstadt". {{cite journal}}: Cite journal requires |journal= (help)
^ abWeitzel, K. (1933). "Amphiperca multiformis ngn sp. und Thaumaturus intermedius n. sp., Knochenfische aus dem Mitteleozän von Messel". Notizblatt des Hessischen Vereins für Erdkunde und der Hessischen Geologischen Landesanstalt zu Darmstadt. 5 (14): 89–97.
^Gaudant, J.; Micklich, N. (1990). "Rhenanoperca minuta nov. gen., nov. sp., ein neuer Percoide (Pisces, Perciformes) aus der Messel-Formation (Mittel-Eozän, Unteres Geiseltalium)". Paläontologische Zeitschrift. 64 (3): 269–286. doi:10.1007/BF02985719. S2CID82717028.
^Micklich, N.; Arratia, G. (2020). "The enigmatic teleostean fish, Thaumaturus intermedius Weitzel, 1933 from the Eocene of Lake Messel (Hessen, S Germany). Part I: Anatomy and taxonomy revised". Palaeontographica Abteilung A. 323 (1–3): 1. Bibcode:2022PalAA.323....1M. doi:10.1127/pala/2022/0125.
^Westphal, F. (1980). "Chelotriton robustus n. sp., ein Salamandride aus dem Eozän der Grube Messel bei Darmstadt". Senckenbergiana Lethaea. 60 (4/6): 475–487.
^Smith, K.T.; Schaal, S.F.K.; Habersetzer, J. (2018). "9 Amphibians in Messel - in the Water and on Land". Messel-An Ancient Greenhouse Ecosystem. Senckenberg. pp. 123–147.
^Roček, Z.; Wuttke, M.; Gardner, J. D.; Singh Bhullar, B.-A. (2014). "The Euro-American genus Eopelobates, and a re-definition of the family Pelobatidae (Amphibia, Anura). Palaeobiodiversity and Palaeoenvironments". 94 (4): 529–567. Bibcode:2014PdPe...94..529R. doi:10.1007/s12549-014-0169-5. S2CID86195679. {{cite journal}}: Cite journal requires |journal= (help)
^Smith, K.T. (2009). "Eocene Lizards of the Clade Geiseltaliellus from Messel and Geiseltal, Germany, and the Early Radiation of Iguanidae (Reptilia: Squamata)". Bulletin of the Peabody Museum of Natural History. 50 (2): 219–306. doi:10.3374/014.050.0201. S2CID84085505.
^Müller, J. (2001). "Osteology and relationships of Eolacerta robusta, a lizard from the Middle Eocene of Germany (Reptilia, Squamata)". Journal of Vertebrate Paleontology. 21 (2): 261–278. doi:10.1671/0272-4634(2001)021[0261:oaroer]2.0.co;2. S2CID86300545.
^Čerňanský, A.; Smith, K. T. (2017). "Eolacertidae: a new extinct clade of lizards from the Palaeogene; with comments on the origin of the dominant European reptile group – Lacertidae". Historical Biology. 30 (7): 994–1014. doi:10.1080/08912963.2017.1327530. S2CID49546941.
^ abcdSmith, K.T.; Čerňanský, A.; Scanferla, A. (2018). "10.1 Lizards and snakes-Warmth-loving sunbathers". Messel-An Ancient Greenhouse Ecosystem. Senckenberg. pp. 123–147.
^ abHervet, S. (2004). "Systématique du groupe "Palaeochelys sensu lato – Mauremys" (Chelonii, Testudinoidea) du Tertiaire d'Europe occidentale: principaux résultats". Annales de Paléontologie. 90 (1): 13–78. doi:10.1016/j.annpal.2003.12.002. ISSN0753-3969.
^Schleich, H. H. (1993). "New reptile material from the German Tertiary. 11. Neochelys franzeni n. sp., the first pleurodiran turtle from Messel". Kaupia. 3: 15–21.
^Delfino, M.; Smith, T. (2012). "Reappraisal of the morphology and phylogenetic relationships of the middle Eocene alligatoroid Diplocynodon deponiae (Frey, Laemmert, and Riess, 1987) based on a three-dimensional specimen". Journal of Vertebrate Paleontology. 32 (6): 1358–1369. doi:10.1080/02724634.2012.699484. S2CID84977303.
^Weitzel, K. (1935). "Hassiacosuchus haupti n. g n. s. ein durophages Krokodil aus dem Mitteleozän von Messel". Notizblatt des Vereins für Erdkunde und der hessischen geologischen Landesanstalt zu Darmstadt. 16: 40–49.
^Rossmann, T.; Rauhe, M.; Ortega, F. (2000). "Studies on Cenozoic crocodiles: 8. Bergisuchus dietrichbergi Kuhn (Sebecosuchia: Bergisuchidae n. fam.) from the Middle Eocene of Germany, some new systematic and biological conclusions". Paläontologische Zeitschrift. 74 (3): 379–392. doi:10.1007/BF02988108. S2CID129507618.
^Brochu, C. A. (2013). "Phylogenetic relationships of Palaeogene ziphodont eusuchians and the status of Pristichampsus Gervais, 1853". Earth and Environmental Science Transactions of the Royal Society of Edinburgh. 103 (3–4): 521–550. doi:10.1017/S1755691013000200. S2CID128920027.
^Rossmann, T. (2002). "Studies on Cenozoic Crocodiles: 10. First Evidence of a Tomistomid Crocodilian (Eusuchia: Tomistomidae) from the Middle Eocene (Geiseltalien, MP11) of Grube Messel, Germany". Neues Jahrb. Geol. Paläontol. Monatsh. 2002 (3): 129–146. doi:10.1127/njgpm/2002/2002/129.
^ abcdMayr, G. (2017). "The early Eocene birds of the Messel fossilsite: a 48 million-year-old bird community adds a temporal perspective to the evolution of tropical avifaunas". Biol. Rev. 92 (2): 1174–1188. doi:10.1111/brv.12274. PMID27062331. S2CID3464510.
^Mayr, G. (2008). "First substantial Middle Eocene record of the Lithornithidae (Aves): A postcranial skeleton from Messel (Germany)". Annales de Paléontologie. 94 (1): 29–37. doi:10.1016/j.annpal.2007.12.004.
^Mayr, G. (2014). "The middle Eocene European "ratite" Palaeotis (Aves, Palaeognathae) restudied once more". Paläontologische Zeitschrift. 89 (3): 503–514. doi:10.1007/s12542-014-0248-y. S2CID83787063.
^Hellmund M (2013). "Reappraisal of the bone inventory of Gastornis geiselensis (Fischer, 1978) from the Eocene Geiseltal Fossillagerstatte (Saxony-Anhalt, Germany)". Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. 269 (2): 203–220. doi:10.1127/0077-7749/2013/0345.
^Angst D.; Lécuyer C.; Amiot R.; Buffetaut E.; Fourel F.; Martineau F.; Legendre S.; Abourachid A.; Herrel A. (2014). "Isotopic and anatomical evidence of an herbivorous diet in the Early Tertiary giant bird Gastornis. Implications for the structure of Paleocene terrestrial ecosystems". Naturwissenschaften. 101 (4): 313–322. Bibcode:2014NW....101..313A. doi:10.1007/s00114-014-1158-2. PMID24563098. S2CID18518649.
^Mayr, Gerald (2006). "New specimens of the early Eocene stem group galliform Paraortygoides (Gallinuloididae), with comments on the evolution of a crop in the stem lineage of Galliformes". Journal of Ornithology. 147 (1): 31–37. Bibcode:2006JOrni.147...31M. doi:10.1007/s10336-005-0006-8. S2CID30402641.
^ abMayr, G. (2016). "Fragmentary but distinctive: three new avian species from the early Eocene of Messel, with the earliest record of medullary bone in a Cenozoic bird". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 279 (3): 273–286. doi:10.1127/njgpa/2016/0555.
^ abMayr, G. (2001). "Comments on the osteology of Masillapodargus longipes Mayr 1999 and Paraprefica major Mayr 1999, caprimulgiform birds from the Middle Eocene of Messel (Hessen, Germany)". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. 2001 (2): 65–76. doi:10.1127/njgpm/2001/2001/65.
^Mayr, G. (2005). "The Palaeogene Old World potooParapreficaMayr, 1999 (Aves, Nyctibiidae): Its osteology and affinities to the New World Preficinae Olson, 1987". Journal of Systematic Palaeontology. 3 (4): 359–370. Bibcode:2005JSPal...3..359M. doi:10.1017/s1477201905001653. S2CID85093005.
^Mayr, G. (2002). "A contribution to the osteology of the Middle Eocene ibis Rhynchaeites messelensis (Aves: Threskiornithidae: Rhynchaeitinae nov. subfam.)". Neues Jahrbuch für Geologie und Paläontologie - Monatshefte. 2002 (8): 501–512. doi:10.1127/njgpm/2002/2002/501.
^Mayr, G. (2005). "A new eocene Chascacocolius-like mousebird (Aves: Coliiformes) with a remarkable gaping adaptation". Organisms Diversity & Evolution. 5 (3): 167–171. Bibcode:2005ODivE...5..167M. doi:10.1016/j.ode.2004.10.013.
^Mayr, G.; Mourer-Chauviré, C. (2000). "Rollers (Aves: Coraciiformes s.s.) from the Middle Eocene of Messel (Germany) and the Upper Eocene of the Quercy (France)". Journal of Vertebrate Paleontology. 20 (3): 533–546. doi:10.1671/0272-4634(2000)020[0533:racssf]2.0.co;2. S2CID56065258.
^ abMayr, G.; Peters, D. S. (1998). "The mousebirds (Aves: Coliiformes) from the Middle Eocene of Grube Messel (Hessen, Germany)". Senckenbergiana Lethaea. 78 (1–2): 179–197. doi:10.1007/bf03042768. S2CID82480792.
^ abMayr, G. (2006). "New specimens of the Eocene Messelirrisoridae (Aves: Bucerotes), with comments on the preservation of uropygial gland waxes in fossil birds from Messel and the phylogenetic affinities of Bucerotes". Paläontol. Z. 80 (4): 390–405. doi:10.1007/BF02990211. S2CID51778381.
^Weidig, I. (2006). "The first New World occurrence of the Eocene birdPlesiocathartes (Aves: ?Leptosomidae)". Paläontologische Zeitschrift. 80 (3): 230–237. doi:10.1007/bf02988439. S2CID140684204.
^Mayr, G.; Mourer-Chauviré, C.; Weidig, I. (2004). "Osteology and systematic position of the eocene primobucconidae (aves, coraciiformes sensu stricto), with first records from Europe". Journal of Systematic Palaeontology. 2 (1): 1–12. Bibcode:2004JSPal...2....1M. doi:10.1017/S1477201903001093. S2CID34962540.
^Mayr, G. (2001). "New specimens of the Middle Eocene fossil mousebird Selmes absurdipes Peters 1999". Ibis. 143 (4): 427–434. doi:10.1111/j.1474-919x.2001.tb04943.x.
^Waterhouse, D. M. (2006). "Parrots in a nutshell: The fossil record of Psittaciformes (Aves)". Historical Biology. 18 (2): 227–238. doi:10.1080/08912960600641224. S2CID83664072.
^ abcdeMayr, G. (2016). "New species of Primozygodactylus from Messel and the ecomorphology and evolutionary significance of early Eocene zygodactylid birds (Aves, Zygodactylidae)". Historical Biology. 29 (7): 875–884. doi:10.1080/08912963.2016.1261135. S2CID89599089.
^Gerald Mayr (2015). "A reassessment of Eocene parrotlike fossils indicates a previously undetected radiation of zygodactyl stem group representatives of passerines (Passeriformes)". Zoologica Scripta. 44 (6): 587–602. doi:10.1111/zsc.12128. S2CID85599482.
^Mayr, G. (2008). "Pumiliornis tessellatus Mayr, 1999 revisited — new data on the osteology and possible phylogenetic affinities of an enigmatic Middle Eocene bird". Paläontologische Zeitschrift. 82 (3): 247–253. doi:10.1007/bf02988891. S2CID55757331.
^Mayr, G. (2016). "Towards completion of the early Eocene aviary: A new bird group from the Messel oil shale (Aves, Eopachypterygidae, fam. nov.)". Zootaxa. 4013 (2): 252–264. doi:10.11646/zootaxa.4013.2.6. PMID26623896.
^Von Koenigswald, W. (1990). "Die Paläobiologie der Apatemyiden (Insectivora sl) und die Ausdeutung der Skelettfunde von Heterohyus nanus aus dem Mitteleozän von Messel bei Darmstadt". Palaeontographica. Abteilung A, Paläozoologie, Stratigraphie. 210 (1–3): 41–77.
^Franzen, J. L. (1988). "Skeletons of Aumelasia (Mammalia, Artiodactyla, Dichobunidae) from Messel (M. Eocene, W. Germany)". Courier Forschungsinstitut Senckenberg. 107: 309–321.
^Franzen, J.L. (1983). "Fossilfundstelle Messel, Nr. 35: Ein zweites Skelett von Messelobunodon (Mammalia, Artiodactyla, Dichobunidae) aus der "Grube Messel" bei Darmstadt (Deutschland, S-Hessen)". Senckenbergiana Lethaea. 64 (5–6): 403–445.
^Tobien, H. (1985). "Zur Osteologie von Masillabune (Mammalia, Artiodactyla, Haplobunodontidae) aus dem Mitteleozan der Fossilfundstatte Messel bei Darmstadt (S-Hessen, Bundesrepublik Deutschland)". Geologisches Jahrbuch Hessen. 113: 5–58.
^Storch, G.; Habersetzer, J. (1988). "Archaeonycteris pollex (Mammalia: Chiroptera) eine neue Fledermaus aus dem Eozän der Grube Messel bei Darmstadt". Courier Forschungsinstitut Senckenberg. 107: 263–273.
^Revilliod, P. (1917). "Contribution à l'étude des chiroptères des terrains tertiares". Abhandlungen der Schweizerischen Paläontologischen Gesellschaft. 42: 1–57.
^ abSmith, J.D.; Storch, G. (1981). "New Middle Eocene bats from "Grube Messel" near Darmstadt, W-Germany". Senckenbergiana Biologica. 61 (3–4): 153–167.
^Storch, G.; Sigé, B.; Habersetzer, J. (2002). "Tachypteron franzeni n. gen., n. sp., earliest emballonurid bat from the Middle Eocene of Messel (Mammalia, Chiroptera)". Paläontol Z. 76 (2): 189–199. Bibcode:2002PalZ...76..189S. doi:10.1007/BF02989856. S2CID85053734.
^Rose, K. D.; Dunn, R. H.; Grande, L. (2014). "A new skeleton ofPalaeosinopa didelphoides(Mammalia, Pantolesta) from the early Eocene Fossil Butte Member, Green River Formation (Wyoming), and skeletal ontogeny in Pantolestidae". Journal of Vertebrate Paleontology. 34 (4): 932–940. Bibcode:2014JVPal..34..932R. doi:10.1080/02724634.2013.822876. S2CID130601648.
^Storch, G. (1993). "Morphologie und Paläobiologie von Macrocranion tenerum, einem Erinaceomorphen aus dem Mittel-Eozän von Messel bei Darmstadt (Mammalia, Lipotyphla)". Senckenbergiana Lethaea. 73 (1): 61–81.
^von Koenigswald, W.; Storch, G. (1983). "Fossilfundstelle Messel, Nr. 36: Pholidocercus hassiacus, ein Amphilemuride aus dem Eozän der "Grube Messel" bei Darmstadt (Mammalia, Lipotyphla)". Senckenbergiana Lethaea. 64 (5–6): 447–495.
^Storch, G.; Lister, A. M. (1985). "Lepticticium nasutum, ein Pseudorhyncocyonide aus dem Eozän der Grube Messel bei Darmstadt (Mammalia, Proteutheria). Fossilfundstelle Messel, Nr. 42". Senckenbergiana Lethaea. 66 (1–2): 1–37.
^Wighart von Koenigswald; Gerhard Storch (March 16, 1987). "Leptictidium tobieni n sp., ein dritter Pseudorhyncocyonide (Proteutheria, Mammalia) aus dem Eozän von Messel". Courier Forschungsinstitut Senckenberg (in German). 91: 107–116, 9 Abb.
^Storch, G. (1993). "Amphiperatherium goethei, ein weiteres Beuteltier aus dem Eozän von Messel (Mammalia, Didelphidae)". Carolinea. 51: 123–124.
^ abSpringhorn, R. (1982). "Neue Raubtiere (Mammalia: Creodonta et Carnivora) aus dem Lutetium der Grube Messel (Deutschland)". Palaeontographica Abteilung A. 179: 105–141.
^Springhorn, R. (2000). "Messelogale, eine neue Raubtiergattung aus dem Mitteleozän von Messel (Deutschland)". Paläontol. Z. 74 (3): 425–439. doi:10.1007/BF02988110. S2CID127809369.
^Springhorn, R. (1980). "Paroodectes feisti, der erste Miacide (Carnivora, Mammalia) aus dem Mittel-Eozän von Messel". Paläontol. Z. 54 (1–2): 171–198. doi:10.1007/BF02985887. S2CID128857759.
^Franzen, J.L. (2006). "Eurohippus n.g., a new genus of horses from the Middle to Late Eocene of Europe". Senckenbergiana Lethaea. 86: 97–102. doi:10.1007/BF03043638. S2CID84192738.
^Franzen, J.L.; Habersetzer, J. (2017). "Complete skeleton of Eurohippus messelensis (Mammalia, Perissodactyla, Equoidea) from the early middle Eocene of Grube Messel (Germany)". Palaeobio Palaeoenv. 97 (4): 807–832. Bibcode:2017PdPe...97..807F. doi:10.1007/s12549-017-0280-5. S2CID135363952.
^Franzen, J. L. (1990). "Hallensia (Mammalia, Perissodactyla) aus Messel und dem Pariser Becken sowie Nachträge aus dem Geiseltal". Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. 60: 175–201.
^Franzen, J. L. (1981). "HYRACHYUS MINIMUS (MAMMALIA, PERISSODACTYLA, HELALETIDAE) AUS DEN MITTELEOZAENEN OELSCHIEFERN DER" GRUBE MESSEL" BEI DARMSTADT (DEUTSCHLAND, S-HESSEN)". Senckenb. Lethaea. 61 (3–6): 371–376. ISSN0037-2110.
^Wilde, V.; Hellmund, M. (2010). "First record of gut contents from a middle Eocene equid from the Geiseltal near Halle (Saale), Sachsen-Anhalt, Central Germany". Palaeobiodiversity and Palaeoenvironments. 90 (2): 153–162. Bibcode:2010PdPe...90..153W. doi:10.1007/s12549-010-0028-y. S2CID128890140.
^Heissig, K. (2003). "Origin and early dispersal of the squirrels and their relatives". Deinsea. 10 (1): 277–286.
^Storch, G.; Seiffert, C. (2007). "Extraordinarily preserved specimen of the oldest known glirid from the middle Eocene of Messel (Rodentia)". Journal of Vertebrate Paleontology. 27 (1): 189–194. doi:10.1671/0272-4634(2007)27[189:EPSOTO]2.0.CO;2. S2CID85909806.