Wangshi Group

Wangshi Group
Wangshi Group
Stratigraphic range: Coniacian-Campanian; ~85–75 Ma PreꞒ Ꞓ O S D C P T J K Pg N
	An outcrop of the Wangshi Group at Kugou which is the type locality for Anomalipes zhaoi. Photographed in c. 2018.
Type	Group
Unit of	Jiaolai Basin
Sub-units	Linjiazhuang, Jiangjunding, Xingezhuang, Hongtuya, Jingangkou & Shijiatun Formations
Underlies	Jiaozhou Formation
Overlies	Qingshan Group
Lithology
Primary	Conglomerate
Other	Marl
Location
Coordinates	36°54′N 120°42′E / 36.9°N 120.7°E
Approximate paleocoordinates	37°18′N 111°00′E / 37.3°N 111.0°E
Region	Shandong
Country	China
	Wangshi Group (China) Wangshi Group (Shandong)

The Wangshi Group (Chinese: 王氏群; pinyin: Wángshì Qún) is a geological Group in Shandong, China whose strata date back to the Coniacian to Campanian stages of the Late Cretaceous.^[1] Dinosaur remains are among the fossils that have been recovered from the group.^[2]

Vertebrate paleofauna

Dinosaurs

Dinosaurs of the Wangshi Group
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Anomalipes^[3]	A. zhaoi			"Partial left hindlimb"	A caenagnathid oviraptorosaur
Chingkankousaurus^[4]	C. fragilis^[4]				Nomen dubium
Ischioceratops^[5]	I. zhuchengensis
Laiyangosaurus^[6]	L. youngi		Jingangkou Formation		A saurolophine hadrosaur
Micropachycephalosaurus^[4]	M. hongtuyanensis^[4]			"Partial mandible, associated postcranial fragments."^[7]
Pinacosaurus^[8]	P. cf. grangeri^[8]	Shandong		"A well-preserved sacrum with the attached right ilium and part of the presacral rod, caudal vertebrae, a left femur and a dermal scute."^[9]	Remains collected in 1923 by H. C. T’an and Otto Zdansky and mentioned by Buffetaut (1995)^[9]
Shantungosaurus^[4]	S. giganteus^[4]		Xingezhuang Formation
Sinoceratops	S. zhuchengensis		Xingezhuang Formation
Tanius^[4]	T. chingkankouensis^[4]		Jingangkou Formation
	T. laiyangensis^[4]		Jingangkou Formation		Nomen dubium
	T. sinensis^[4]		Jingangkou Formation
Tsintaosaurus^[4]	T. spinorhinus^[4]		Jingangkou Formation	"Isolated skull and postcranial elements from at least [five] individuals."^[10]
Zhuchengtyrannus^[11]	Z. magnus^[11]		Xingezhuang Formation
Zhuchengceratops	Z. inexpectus		Xingezhuang Formation
Zhuchengtitan^[12]	Z. zangjiazhuangensis		Xingezhuang Formation	"a single humerus"	A titanosaur closely related to Opisthocoelicaudia

Other fauna

Other fauna of the Wangshi Group
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Shandongemys^[13]	S. dongwuica				A lindholmemydid turtle

Fossil eggs

The following fossil eggs were recovered from the Jingangkou Formation of the Wangshi Group.^[14]^[15]

Saurischia
- Elongatoolithidae
  - Elongatoolithus elongatus
Neornithischia
- Ovaloolithidae
  - Ovaloolithus chinkangkouensis
  - Ovaloolithus laminadermus

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

References

^ Wangshi Group in the Paleobiology Database
^ Weishampel et al., 2004, pp.593-600
^ Yilun Yu; Kebai Wang; Shuqing Chen; Corwin Sullivan; Shuo Wang; Peiye Wang; Xing Xu (2018). "A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with comments on size variation among oviraptorosaurs". Scientific Reports. 8: Article number 5030. doi:10.1038/s41598-018-23252-2
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l "48.9 Shandong, People's Republic of China; 1. Wangshi Group," in Weishampel, et al. (2004). Page 598.
^ Yiming He, Peter J. Makovicky, Kebai Wang, Shuqing Chen, Corwin Sullivan, Fenglu Han, Xing XuMichael J. Ryan, David C. Evans, Philip J. Currie, Caleb M. Brown and Don Brinkman (2015). "A New Leptoceratopsid (Ornithischia, Ceratopsia) with a Unique Ischium from the Upper Cretaceous of Shandong Province, China".
^ Zhang, J.L.; Wang, X.; Wang, Q.; Jiang, S.; Cheng, X.; Ning, L.; Qiu, R. (2017). "A new saurolophine hadrosaurid (Dinosauria: Ornithopoda) from the Upper Cretaceous of Shandong, China" (PDF). Annals of the Brazilian Academy of Sciences.
^ "Table 21.1," in Weishampel, et al. (2004). Page 466.
^ ^a ^b Listed as "Pinacosaurus cf. grangeri" in "48.9 Shandong, People's Republic of China; 1. Wangshi Group," in Weishampel, et al. (2004). Page 598.
^ ^a ^b Buffetaut, Eric (1995). "An ankylosaurid dinosaur from the Upper Cretaceous of Shandong (China)". Geological Magazine. 132 (6): 683–692. Bibcode:1995GeoM..132..683B. doi:10.1017/s0016756800018914. ISSN 0016-7568.
^ "Table 20.1," in Weishampel, et al. (2004). Page 442.
^ ^a ^b David W. E. Hone; Kebai Wang; Corwin Sullivan; Xijin Zhao; Shuqing Chen; Dunjin Li; Shuan Ji; Qiang Ji; Xing Xu (2011). "A new, large tyrannosaurine theropod from the Upper Cretaceous of China". Cretaceous Research. 32 (4): 495–503. Bibcode:2011CrRes..32..495H. doi:10.1016/j.cretres.2011.03.005.
^ Mo, J.; Wang, K.; Chen, S.; Wang, P.; Xu, X. (2017). "A new titanosaurian sauropod from the Late Cretaceous strata of Shandong Province". Geological Bulletin of China. 36 (9): 1501–1505.
^ L. Li, H. Tong, K. Wang, S. Chen, and X. Xu. 2013. Lindholmemydid turtles (Cryptodira: Testudinoidea) from the Late Cretaceous of Shandong Province, China. Annales de Paléontologie 99:243-259
^ Jingangkou in the Paleobiology Database
^ Young, 1954

Bibliography

Weishampel, David B.; Dodson, Peter; Osmólska, Halszka, eds. (2004). The Dinosauria, 2nd edition. Berkeley: University of California Press. ISBN 0-520-24209-2. Retrieved 2019-02-21.
Hu, Cheng, Pang and Fang (2001). Shantungosaurus giganteus: [3 front matter] + ii + 139 pp. + 18 plates (in Chinese with English abstract; publisher's name not translated) ISBN 7-116-03472-2
Young, C.-C (1954). "Fossil reptilian eggs from Laiyang, Shantung, China". Scientia Sinica. 3: 505–522.

[PBDB-1] Wangshi Group in the Paleobiology Database

[cretaceousdistribution-2] Weishampel et al., 2004, pp.593-600

[3] Yilun Yu; Kebai Wang; Shuqing Chen; Corwin Sullivan; Shuo Wang; Peiye Wang; Xing Xu (2018). "A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with comments on size variation among oviraptorosaurs". Scientific Reports. 8: Article number 5030. doi:10.1038/s41598-018-23252-2

[wangshi-4] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l "48.9 Shandong, People's Republic of China; 1. Wangshi Group," in Weishampel, et al. (2004). Page 598.

[5] Yiming He, Peter J. Makovicky, Kebai Wang, Shuqing Chen, Corwin Sullivan, Fenglu Han, Xing XuMichael J. Ryan, David C. Evans, Philip J. Currie, Caleb M. Brown and Don Brinkman (2015). "A New Leptoceratopsid (Ornithischia, Ceratopsia) with a Unique Ischium from the Upper Cretaceous of Shandong Province, China".

[6] Zhang, J.L.; Wang, X.; Wang, Q.; Jiang, S.; Cheng, X.; Ning, L.; Qiu, R. (2017). "A new saurolophine hadrosaurid (Dinosauria: Ornithopoda) from the Upper Cretaceous of Shandong, China" (PDF). Annals of the Brazilian Academy of Sciences.

[table-21-1-466-7] "Table 21.1," in Weishampel, et al. (2004). Page 466.

[pinaco-8] Listed as "Pinacosaurus cf. grangeri" in "48.9 Shandong, People's Republic of China; 1. Wangshi Group," in Weishampel, et al. (2004). Page 598.

[:0-9] Buffetaut, Eric (1995). "An ankylosaurid dinosaur from the Upper Cretaceous of Shandong (China)". Geological Magazine. 132 (6): 683–692. Bibcode:1995GeoM..132..683B. doi:10.1017/s0016756800018914. ISSN 0016-7568.

[table-20-1-442-10] "Table 20.1," in Weishampel, et al. (2004). Page 442.

[Zhuchengtyrannus-11] David W. E. Hone; Kebai Wang; Corwin Sullivan; Xijin Zhao; Shuqing Chen; Dunjin Li; Shuan Ji; Qiang Ji; Xing Xu (2011). "A new, large tyrannosaurine theropod from the Upper Cretaceous of China". Cretaceous Research. 32 (4): 495–503. Bibcode:2011CrRes..32..495H. doi:10.1016/j.cretres.2011.03.005.

[12] Mo, J.; Wang, K.; Chen, S.; Wang, P.; Xu, X. (2017). "A new titanosaurian sauropod from the Late Cretaceous strata of Shandong Province". Geological Bulletin of China. 36 (9): 1501–1505.

[13] L. Li, H. Tong, K. Wang, S. Chen, and X. Xu. 2013. Lindholmemydid turtles (Cryptodira: Testudinoidea) from the Late Cretaceous of Shandong Province, China. Annales de Paléontologie 99:243-259

[PBDBJingangkou-14] Jingangkou in the Paleobiology Database

[Young1954-15] Young, 1954

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