A member of Galeaspida. Genus includes new species Q. elaia.
Jawless fishes research
A study on the phylogenetic relationships of cyathaspidids is published by Elliott, Lassiter & Blieck (2021).[5]
Miyashita et al. (2021) report larval and juvenile forms of four stem lampreys from the Paleozoic era (Hardistiella, Mayomyzon, Pipiscius and Priscomyzon), including a hatchling-to-adult growth series of Priscomyzon, and report that the studied larvae display features that are otherwise unique to adult modern lampreys, and lack the defining traits of ammocoetes.[6]
A study on the anatomy and likely feeding ecology of Mesomyzon mengae, based on data from new, well-preserved specimens, is published by Wu, Chang & Janvier (2021).[7]
A study on the histology of the dermal skeleton in Procephalaspis oeselensis, Aestiaspis viitaensis, Dartmuthia gemmifera and four species of Tremataspis is published by Bremer et al. (2021), who interpret their findings as indicative of the emergence of the complex pore-canal system in Tremataspis through the modification of the structures already present in other taxa.[8]
A study aiming to determine whether the earliest vertebrates may have swum under various conditions without a clearly-differentiated tail fin, based on data from an abstracted model of Metaspriggina walcotti, is published by Rival, Yang & Caron (2021).[9]
A study on the morphological and functional diversity of osteostracan and galeaspid headshields, and on its implications for the knowledge of the ecology of the immediate jawless relatives of jawed vertebrates, is published by Ferrón et al. (2021).[10]
Redescription of Nochelaspis maeandrine is published by Meng, Zhu & Gai (2021).[11]
A study on the anatomy of a dorsal head shield of Kalanaspis delectabilis is published by Tinn et al. (2021).[12]
A member of Arthrodira belonging to the family Plourdosteidae. The type species is L. ziregensis.
Placoderm research
Zhu et al. (2021) use CT scanning to reveal the endocast of Brindabellaspis stensioi, and evaluate the implications of its anatomy for the knowledge of the phylogenetic relationships of early jawed vertebrates.[15]
Redescription of the anatomy of the headshield of Parayunnanolepis xitunensis is published by Wang & Zhu (2021).[16]
Description of new fossil material of Palaeacanthaspis vasta from the Devonian (Lochkovian) Chortkiv Formation (Ukraine), and a study on the phylogenetic relationships of this species, is published by Dupret et al. (2021).[17]
A study on the development of teeth in acanthodians, and on its implications for the knowledge of the evolution of teeth of jawed vertebrates, is published by Rücklin et al. (2021).[20]
A study on the anatomy of teeth, jaws and associated oral structures of acanthodians, and on their implications for the knowledge of the evolution of dentition of modern cartilaginous fishes, is published by Dearden & Giles (2021).[21]
A probable planktivorous shark placed in the new family Aquilolamnidae, of uncertain placement. Possibly a member of Lamniformes. The type species is A. milarcae.
A member of the family Jalodontidae. Genus includes "Phoebodus" brodiei Woodward (1893) (interpreted by Ivanov, Duffin & Richter, 2021 as a senior synonym of "Phoebodus" keuperinus Seilacher, 1948).
A member of Neoselachii; a replacement name for Vallisia Duffin (1982).
Cartilaginous fish research
Description of new fossil material of Gualepis elegans from the Lower Devonian of Yunnan (China), and a study on the phylogenetic relationships of this fish, is published by Cui et al. (2021).[46]
Mottequin et al. (2021) reject the interpretation of Spiraxis interstrialis as chondrichthyan egg cases, and evaluate the implications of this reinterpretation for the knowledge of the evolution of oviparity in cartilaginous fishes.[47]
Description of the first known skull remains of Onchopristis numidus from the Cretaceous Kem Kem Group (Morocco), and a study on the anatomy and phylogenetic relationships of this species, is published by Villalobos-Segura et al. (2021), who name a new family Onchopristidae.[48]
New, exceptionally well-preserved skeleton of Asteracanthus ornatissimus, preserved with teeth that markedly differ from other teeth referred to Asteracanthus, is described from the TithonianAltmühltal Formation (Germany) by Stumpf et al. (2021), who interpret this specimen as indicating that Asteracanthus and Strophodus represent two valid genera distinct from all other hybodontiforms.[49]
A study on the morphological diversity of teeth of lamniform sharks from mid-Cretaceous assemblages in Australia, and on its implications for the knowledge of the composition of mid-Cretaceous shark communities and their recovery in the aftermath of the Cenomanian-Turonian boundary event, is published by Bazzi, Kear & Siversson (2021).[50]
A study on the biomechanics of teeth of five species of Otodus, aiming to assess the functional significance of morphological trends in otodontid teeth and to test whether the morphology of otodontid teeth enabled the transition from piscivory to predation on marine mammals and the evolution of titanic body sizes, is published by Ballell & Ferrón (2021)[51]
A study on growth patterns, reproductive biology and likely lifespan of Otodus megalodon is published by Shimada et al. (2021).[53]
Perez, Leder & Badaut (2021) present a novel method for estimating body size in fossil lamniform sharks, and attempt to determine the body size of Otodus megalodon.[54]
Revision of the fossil record of the extant tiger shark and the extinct members of the tiger shark lineage is published by Türtscher et al. (2021).[55]
Redescription of Striatolamia tchelkarnurensis is published by Malyshkina (2021).[56]
Shark teeth which might represent the first occurrence of the blacknose shark in the Pacific Ocean are described from the Pliocene Upper Onzole Formation (Ecuador) by Collareta et al. (2021), who evaluate the implications of this finding for the knowledge of the evolutionary history of the blacknose shark and the whitenose shark.[57]
Two fossil teeth of the blacktip shark are reported from lower Pliocene marine deposits of Tuscany (Italy) by Collareta et al. (2021), representing the first known occurrence of this species in the fossil record from both Europe and the Mediterranean Basin.[58]
A study on the morphological diversity of extant and fossil shark teeth, and on their implications for the knowledge of the evolution of lamniform and carcharhiniform sharks throughout the last 83 million years, is published by Bazzi et al. (2021).[59]
A study on the evolutionary history of sharks across the Cretaceous–Paleogene extinction event, as indicated by morphological diversity of shark teeth across the Cretaceous–Paleogene interval, is published by Bazzi et al. (2021).[60]
Evidence of a previously unknown major extinction of sharks in the early Miocene, ~19 million years ago, is presented by Sibert & Rubin (2021);[61] the study is subsequently criticized by Naylor et al. (2021)[62][63] and Feichtinger et al. (2021).[64][65]
A study on shark scales from mid-Holocene (~7,000-y-old) and modern reef sediments in Bocas del Toro (Panama), aiming to determine changes of shark abundance in this area since the mid-Holocene and their possible causes, is published by Dillon et al. (2021).[66]
A member of the subfamily Gobionellinae. The type species is H. praeschismatus; genus also includes "Pomatoschistus" bunyatovi Bratishko et al. (2015).
A member of the family Gobiidae. The type species is S. compactus; genus also includes "Hesperichthys" iugosus Schwarzhans, Brzobohatý & Radwańska (2020).
A member of the family Gobiidae. The type species is Y. decoratus; genus also includes Y. naslavcensis.
Ray-finned fish research
Putative paraxial ossicle of a member of the family Molidae, possibly representing the first fossil find of the genus Mola from the Mediterranean Basin reported to date, is described from the Miocene Pietra Leccese Formation (Apulia, Italy) by Collareta et al. (2021).[111]
A platysomid specimen, representing the earliest deep-bodied actinopterygian reported to date, is described from the Carboniferous (Tournaisian) Horton Bluff Formation (Canada) by Wilson, Mansky & Anderson (2021), who evaluate the implications of this findings for the knowledge of the evolution of early ray-finned fishes.[112]
A review of the fossil record of Early–Middle Triassic marine bony fishes, aiming to determine the implications of poor fossil record from the late Olenekian-early middle Anisian interval on the knowledge of the Triassic radiation of bony fishes, is published by Romano (2021).[113]
New fish fauna dating to the Paleocene–Eocene Thermal Maximum, indicating that diverse fish communities thrived in the paleotropics during this time period, is reported from Egypt by El-Sayed et al. (2021).[116]
Heingård et al. (2021) report preservation of residues of both internal and integumentary tissues in the form of dark organic stains in fossil fish larvae from the Eocene (Ypresian) Stolleklint Clay (Ølst Formation, Denmark).[117]
Revision of the fossil material of sturgeons from the Upper Miocene deposits of southern Ukraine is published by Hilton, Kovalchuk & Podoplelova (2021).[118]
A study on the morphological diversity and evolution of pycnodontiforms is published by Cawley et al. (2021).[119]
A study on fossil crushing dentitions of Pycnodus zeaformis and P. maliensis, providing evidence of a distinct pattern of gap-filling tooth addition in pycnodonts, with individual large teeth replaced by multiple small teeth, is published by Collins & Underwood (2021).[120]
A redescription of Atacamichthys greeni is published by Arratia et al. (2021), who interpret it as a stem-groupteleost, and name the new family Atacamichthyidae.[122]
A study on genome size evolution in fossil stem-group teleosts (based on data from bone cell volumes in fossil specimens), aiming to determine the timing of whole-genome duplication in the evolutionary history of teleosts, is published by Davesne et al. (2021).[124]
New fossil material of elopomorphs, including the earliest records of members of the genera Albula and Paralbula from Gondwana reported to date and one of the earliest records of the genus Egertonia, is described from the Upper Cretaceous Mahajanga Basin (Madagascar) by Ostrowski (2021).[125]
A study on the evolutionary history of lanternfishes, primarily based on the fossil record of otoliths, is published by Schwarzhans & Carnevale (2021).[126]
A coelacanth specimen belonging or related to the species Heptanema paradoxum is described from the Ladinian Meride Limestone (Switzerland) by Renesto, Magnani & Stockar (2021), representing the first known coelacanth specimen from the Middle Triassic that undoubtedly bears elongate thin ribs.[130]
Fossil material of mawsoniid coelacanths is described from the marine Rhaetian Bonenburg locality (Germany) by Hartung et al. (2021), who interpret this finding as indicating that mawsoniids were already present in Europe in the Late Triassic, and that they inhabited marine environments at the end of the Triassic.[131]
Fossil material of a member of genus Mawsonia is described from the CenomanianWoodbine Formation (Texas, United States; representing the first Cretaceous North American mawsoniid coelacanth reported to date) by Cavin et al. (2021), who evaluate the implications of this finding for the knowledge of potential factors that might have made long survival of the genera Mawsonia and Latimeria possible.[132]
An ossified lung of a mawsoniid coelacanth is described from the Maastrichtian of Oued Zem (Morocco) by Brito et al. (2021), representing the last known record of a Mesozoic coelacanth and the first known occurrence of coelacanths in the phosphate deposits of North Africa.[133]
A study on the evolution of feeding modes among tetrapodomorphs, as indicated by the anatomy of the skull of Tiktaalik roseae, is published by Lemberg, Daeschler & Shubin (2021), who report the simultaneous occurrence of anatomical modifications of the skull for prey capture through biting, as well as joint morphologies suggestive of cranial kinesis that is also present in suction-feeding fish.[134]
A study on the phylogenetic relationships of extant and fossil coelacanths is published by Toriño, Soto & Perea (2021).[135]
A study on the morphology and histology of the scales of Miguashaia bureaui, and on its implications for the knowledge of the evolution of the squamation in coelacanths, is published by Mondéjar-Fernández et al. (2021).[136]
New fossil remains representing one of the largest known coelacanths ever reported are described from the Middle Jurassic of Normandy (France) by Cavin et al. (2021), who also compare the relationship between taxic diversity and body size diversity in coelacanths and ray-finned fishes over the Devonian–Paleocene time interval.[137]
A study on tooth development in Powichthys thorsteinssoni, evaluating its implications for the knowledge of the evolution of the dentition of bony fishes, is published by King, Marone & Rücklin (2021).[138]
A study on the anatomy and phylogenetic relationships of Cladarosymblema narrienense is published by Clement et al. (2021).[139]
General research
A study on the morphology of the earliest osteocytes in Tremataspis mammillata and Bothriolepis trautscholdi is published by Haridyet al. (2021), who interpret their findings as indicating that the earliest known osteocytes in the fossil record had similar morphology and likely similar physiological capabilities to their modern counterparts, and attempt to determine initial driver favoring evolution of cellular (osteocytic) over acellular (anosteocytic) bones in vertebrates.[140]
Two Permian fish assemblages consisting of cartilaginous fishes and ray-finned fishes are reported from the Madumabisa Mudstone Formation (Zambia) by Peecook et al. (2021), who compare these assemblages with middle and late Permian freshwater fish faunas from Australia, Brazil, Chile and South Africa.[141]
A middle Miocene freshwater fish fossil fauna is described from the Castilletes Formation (Colombia) by Ballen et al. (2021), report the presence of members of fish groups known from extant Amazonian faunas east of the Andes but absent from faunas west of the Andes, and interpret their presence as evidence that the riverine systems of the Guajira Peninsula were connected to Amazonia during the middle Miocene.[142]
References
^Meng, X.; Gai, Z. (2021). "Falxcornus, a new genus of Tridensaspidae (Galeaspida, stem-Gnathostomata) from the Lower Devonian in Qujing, Yunnan, China". Historical Biology: An International Journal of Paleobiology. 34 (5): 897–906. doi:10.1080/08912963.2021.1952198. S2CID237702255.
^Liu, Y.-L.; Huang, L.-B.; Zong, R.-W.; Gong, Y.-M. (2021). "The oldest eugaleaspiform (Galeaspida) from the Silurian Fentou Formation (Telychian, Llandovery) of Wuhan, South China". Journal of Systematic Palaeontology. 19 (4): 253–264. Bibcode:2021JSPal..19..253L. doi:10.1080/14772019.2021.1883755. S2CID233647126.
^Jiang, W.; Zhu, M.; Shi, X.; Li, Q.; Gai, Z. (2021). "Qushiaspis, a new genus of gantarostrataspid fish (Galeaspida, stem-Gnathostomata) from the Lower Devonian of Yunnan, China". Historical Biology: An International Journal of Paleobiology. 33 (12): 3714–3722. Bibcode:2021HBio...33.3714J. doi:10.1080/08912963.2021.1888086. S2CID233887595.
^Meng, X.-Y.; Zhu, M.; Gai, Z.-K. (2021). "Redescription of Nochelaspis maeandrine, the largest eugaleaspiform from the Lower Devonian of Qujing, Yunnan". Vertebrata PalAsiatica. 59 (4): 257–272. doi:10.19615/j.cnki.2096-9899.210727.
^Wang, Y.; Zhu, M. (2021). "New data on the headshield of Parayunnanolepis xitunensis (Placodermi, Antiarcha), with comments on nasal capsules in antiarchs". Journal of Vertebrate Paleontology. 40 (6): e1855189. doi:10.1080/02724634.2020.1855189. S2CID233931146.
^ abcCanevet, J.-M.; Lebrun, P. (2021). "Des dents de requins fossiles. 10. Nouvelles espèces de Carcharhinidae du Miocène de l'Ouest de la France". Fossiles. Revue française de paléontologie. 46: 5–22.
^Cicimurri, D. J.; Ebersole, J. A. (2021). "New Paleogene elasmobranch (Chondrichthyes) records from the Gulf Coastal Plain of the United States, including a new species of Carcharhinus de Blainville, 1816". Cainozoic Research. 21 (2): 147–164.
^Feichtinger, I.; Ivanov, A. O.; Winkler, V.; Dojen, C.; Kindlimann, R.; Kriwet, J.; Pfaff, C.; Schraut, G.; Stumpf, S. (2021). "Scarce ctenacanthiform sharks from the Mississippian of Austria with an analysis of Carboniferous elasmobranch diversity in response to climatic and environmental changes". Journal of Vertebrate Paleontology. 41 (2): e1925902. Bibcode:2021JVPal..41E5902F. doi:10.1080/02724634.2021.1925902. S2CID237518044.
^Roelofs, B.; Königshof, P.; Trinajstic, K.; Munkhjargal, A. (2021). "Vertebrate microremains from the Late Devonian (Famennian) of western Mongolia". Palaeobiodiversity and Palaeoenvironments. 101 (3): 741–753. Bibcode:2021PdPe..101..741R. doi:10.1007/s12549-021-00503-1. S2CID237367224.
^Bronson, A. W. (2021). "A three-dimensionally preserved stethacanthid cranium and endocast from the Late Mississippian Fayetteville Shale (Arkansas, USA)". In Pradel, A.; Denton, J. S. S.; Janvier, P. (eds.). Ancient Fishes and their Living Relatives: a Tribute to John G. Maisey. Munich, Germany: Verlag Dr. Friedrich Pfeil. pp. 93–100. ISBN978-3-89937-269-4.
^Long, J. A.; Thomson, V.; Burrow, C. J.; Turner, S. (2021). "Fossil chondrichthyan remains from the Middle Devonian Kevington Creek Formation, South Blue Range, Victoria". In Pradel, A.; Denton, J. S. S.; Janvier, P. (eds.). Ancient Fishes and their Living Relatives: a Tribute to John G. Maisey. Munich, Germany: Verlag Dr. Friedrich Pfeil. pp. 239–245. ISBN978-3-89937-269-4.
^Lebedev, O. A.; Popov, E. V.; Bagirov, S. V.; Bolshiyanov, I. P.; Kadyrov, R. I.; Statsenko, E. O. (2021). "The earliest chimaeriform fish from the Carboniferous of Central Russia". Journal of Systematic Palaeontology. 19 (12): 821–846. Bibcode:2021JSPal..19..821L. doi:10.1080/14772019.2021.1977732. S2CID239509836.
^Sharma, A.; Singh, S. (2021). "A small assemblage of marine hybodont sharks from the Bathonian of the Jaisalmer Basin, India". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 301 (3): 317–333. doi:10.1127/njgpa/2021/1014. S2CID239669413.
^Kumar, K.; Bajpai, S.; Pandey, P.; Ghosh, T.; Bhattacharya, D. (2021). "Hybodont sharks from the Jurassic of Jaisalmer, western India". Historical Biology: An International Journal of Paleobiology. 34 (6): 953–963. doi:10.1080/08912963.2021.1954920. S2CID238781606.
^Luccisano, V.; Pradel, A.; Amiot, R.; Gand, G.; Steyer, J.-S.; Cuny, G. (2021). "A new Triodus shark species (Xenacanthidae, Xenacanthiformes) from the lowermost Permian of France and its paleobiogeographic implications". Journal of Vertebrate Paleontology. 41 (2): e1926470. Bibcode:2021JVPal..41E6470L. doi:10.1080/02724634.2021.1926470. ISSN0272-4634. S2CID237518013.
^Duffin, C. J. (2021). "A replacement name for the preoccupied genus name Vallisia Duffin, 1982 (Chondrichthyes: Neoselachii)". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 301 (2): 229–232. doi:10.1127/njgpa/2021/1005. S2CID238737779.
^Cui, X.; Qu, Q.; Andreev, P. S.; Li, Q.; Mai, H.; Zhu, M. (2021). "Modeling scale morphogenesis in a Devonian chondrichthyan and scale growth patterns in crown gnathostomes". Journal of Vertebrate Paleontology. 41 (2): e1930018. Bibcode:2021JVPal..41E0018C. doi:10.1080/02724634.2021.1930018. S2CID237517966.
^Mottequin, B.; Goolaerts, S.; Hunt, A. P.; Olive, S. (2021). "The erroneous chondrichthyan egg case assignments from the Devonian: implications for the knowledge on the evolution of the reproductive strategy within chondrichthyans". The Science of Nature. 108 (5): Article number 36. Bibcode:2021SciNa.108...36M. doi:10.1007/s00114-021-01751-z. PMID34432151. S2CID237295117.
^Villalobos-Segura, Y.; Kriwet, J.; Vullo, R.; Stumpf, S.; Ward, D. J.; Underwood, C. J. (2021). "The skeletal remains of the euryhaline sclerorhynchoid †Onchopristis (Elasmobranchii) from the 'Mid'-Cretaceous and their palaeontological implications". Zoological Journal of the Linnean Society. 193 (2): 746–771. doi:10.1093/zoolinnean/zlaa166.
^Collareta, A.; Landini, W.; Bianucci, G.; Di Celma, C. (2021). "Until Panama do us part: new finds from the Pliocene of Ecuador provide insights into the origin and palaeobiogeographic history of the extant requiem sharks Carcharhinus acronotus and Nasolamia velox". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 300 (1): 103–115. doi:10.1127/njgpa/2021/0981. S2CID234849316.
^Collareta, A.; Merella, M.; Casati, S.; Di Cencio, A. (2021). "First fossils of the extant blacktip shark Carcharhinus limbatus from Europe and the Mediterranean Basin". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 301 (1): 109–118. doi:10.1127/njgpa/2021/1002. hdl:11568/1117115. ISSN0077-7749. S2CID237731617.
^Naylor, G. J. P.; de Lima, A.; Castro, J. I.; Hubbell, G.; de Pinna, M. C. C. (2021). "Comment on "An early Miocene extinction in pelagic sharks"". Science. 374 (6573): eabj8723. doi:10.1126/science.abj8723. PMID34882454. S2CID245012652.
^Capasso, L.; Tantawy, A. A.; Mousa, M. K.; Wahba, D. G. A.; Abu El-Kheir, G. A. (2021). "Anomoeodus aegypticus n. sp. (Pisces, †Pycnodontiformes) from the late Cretaceous of the Dakhla Formation, Western Desert, Egypt". Thalassia Salentina. 43: 89–104. doi:10.1285/i15910725v43p89.
^Nam, G.; Nazarkin, M. V.; Bannikov, A. F. (2021). "First discovery of the genus Auxis (Actinopterygii: Scombridae) in the Neogene of South Korea". Bollettino della Società Paleontologica Italiana. 60 (1): 61–67. doi:10.4435/BSPI.2021.05 (inactive 2024-11-20).{{cite journal}}: CS1 maint: DOI inactive as of November 2024 (link)
^Hacker, R. J.; Shimada, K. (2021). "A new ichthyodectiform fish (Actinopterygii: Teleostei) from the Arlington Member (mid-Cenomanian) of the Upper Cretaceous Woodbine Formation in Texas, USA". Cretaceous Research. 123: Article 104798. Bibcode:2021CrRes.12304798H. doi:10.1016/j.cretres.2021.104798. S2CID233806833.
^Cantalice, K. M.; Than-Marchese, B. A.; Villalobos-Segura, E. (2021). "A new Cenomanian acanthomorph fish from the El Chango quarry (Chiapas, south-eastern Mexico) and its implications for the early diversification and evolutionary trends of acanthopterygians". Papers in Palaeontology. 7 (3): 1699–1726. Bibcode:2021PPal....7.1699C. doi:10.1002/spp2.1359. S2CID234805660.
^Murray, A. M.; Holmes, R. B. (2021). "A new species of claroteid catfish (Siluriformes: Claroteidae) from the Eocene of Egypt, (Africa) indicates continental differences in tempo of catfish evolution". Journal of Vertebrate Paleontology. 41 (3): e1979021. Bibcode:2021JVPal..41E9021M. doi:10.1080/02724634.2021.1979021. S2CID239508127.
^ abcdSu, R.; Chang, M.-M.; Chen, G. (2021). "Fossil pharyngeal teeth of grass carp group and their implications for evolution, temporal and spatial distribution pattern, and paleoenvironment". Science China Earth Sciences. 64 (11): 1844–1859. Bibcode:2021ScChD..64.1844S. doi:10.1007/s11430-020-9724-7. S2CID237245324.
^Abu El-Kheir, G. A.; Tantawy, A. A.; Mousa, M. K.; Wahba, D. G. A.; Capasso, L. (2022). "Diastemapycnodus tavernensis gen. et sp. nov. (Actinopterygii, †Pycnodontiformes) from the marine Maastrichtian (Late Cretaceous) of the Dakhla Formation, Western Desert, Egypt". Historical Biology: An International Journal of Paleobiology. 34 (12): 2324–2331. Bibcode:2022HBio...34.2324A. doi:10.1080/08912963.2021.2014482. S2CID245051336.
^Ebert, M. (2021). "A new Ophiopsiformes (Halecomorphi, Neopterygii) from the Upper Jurassic of Nusplingen (Germany) and a comparison of Kimmeridgian Ophiopsiformes from Nusplingen with Tithonian taxa from the Solnhofen Archipelago". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 302 (3): 299–329. doi:10.1127/njgpa/2021/1033. S2CID245092175.
^ abWick, S. L. (2021). "New early Campanian characiform fishes (Otophysi: Characiformes) from west Texas support a South American origin for known Late Cretaceous characiforms from North America". Cretaceous Research. 128: Article 104993. Bibcode:2021CrRes.12804993W. doi:10.1016/j.cretres.2021.104993.
^ abSchwarzhans, W.; Agiadi, K.; Thivaiou, D. (2021). "Teleost otoliths from the Aquitanian (early Miocene) of the Felli section in Greece: the roots of the Mediterranean goby stock (Gobiidae, Gobiiformes)". Rivista Italiana di Paleontologia e Stratigrafia. 127 (3): 485–495. doi:10.13130/2039-4942/16126.
^ abcSchultze, H.-P.; Mickle, K. E.; Poplin, C.; Hilton, E. J.; Grande, L., eds. (2021). Actinopterygii I. Palaeoniscimorpha, Stem Neopterygii, Chondrostei. Handbook of Paleoichthyology. Verlag Dr. Friedrich Pfeil. pp. 1–299. ISBN978-3-89937-272-4.
^Přikryl, T. (2021). "Krumvirichthys brzobohatyi gen. et sp. nov. – the oldest record of the deep-sea smelts (Bathylagidae, Argentiniformes)". Rivista Italiana di Paleontologia e Stratigrafia. 127 (3): 585–594. doi:10.13130/2039-4942/16421.
^Nazarkin, M. V.; Bannikov, A. F. (2021). "Portentosoceros, the new genus for the Neogene fish Pentaceros sakhaliniсus Gretchina (Perciformes, Percoidei) from the western Pacific and its taxonomic position". Journal of Vertebrate Paleontology. 41 (2): e1928682. Bibcode:2021JVPal..41E8682N. doi:10.1080/02724634.2021.1928682. S2CID237139110.
^Ren, Y.; Xu, G.-H. (2021). "A new species of Pteronisculus from the Middle Triassic (Anisian) of Luoping, Yunnan, China, and phylogenetic relationships of early actinopterygian fishes". Vertebrata PalAsiatica. 59 (3): 169–199. doi:10.19615/j.cnki.2096-9899.210518. S2CID238786196.
^Yabumoto, Y.; Milàn, M. V. (2021). "A New Miocene Scorpaenoid Fish, Raususetarches sakurai gen. et sp. nov. (Teleostei: Scorpaeniformes) from Rausu, Hokkaido, Japan". Paleontological Research. 25 (2): 93–104. doi:10.2517/2020PR013. ISSN1342-8144. S2CID233029541.
^Renesto, S.; Magnani, F.; Stockar, R. (2021). "A new species of Saurichthys (Actinopterygii: Saurichtydae) from the Middle Triassic of Monte San Giorgio". Rivista Italiana di Paleontologia e Stratigrafia. 127 (1): 49–71. doi:10.13130/2039-4942/15143.
^Stringer, G.; Schwarzhans, W. (2021). "Corrigendum to "Upper Cretaceous teleostean otoliths from the Severn Formation(Maastrichtian) of Maryland, USA, with an unusual occurrence of Siluriformes and Beryciformes and the oldest Atlantic coast Gadiformes"; Cretaceous Research, 125 (2021), 104867, pages 1–29". Cretaceous Research. 128: Article 104939. Bibcode:2021CrRes.12804939S. doi:10.1016/j.cretres.2021.104939. S2CID237812351.
^Collareta, A.; Peri, E.; Carnevale, G.; Bosselaers, M.; Bianucci, G. (2021). "On the presence of an ocean sunfish (Tetraodontiformes, Molidae) in the Miocene Pietra Leccese formation of Southern Italy". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 301 (2): 147–155. doi:10.1127/njgpa/2021/1003. S2CID238694036.
^Sinha, S.; Brinkman, D. B.; Murray, A. M.; Krause, D. W. (2021). "Late Paleocene fishes of the Ravenscrag Formation, Roche Percée area, southeastern Saskatchewan, Canada". Journal of Vertebrate Paleontology. 41 (3): e1957907. Bibcode:2021JVPal..41E7907S. doi:10.1080/02724634.2021.1957907. S2CID244383384.
^El-Sayed, S.; Friedman, M.; Anan, T.; Faris, M. A.; Sallam, H. (2021). "Diverse marine fish assemblages inhabited the paleotropics during the Paleocene-Eocene thermal maximum". Geology. 49 (8): 993–998. Bibcode:2021Geo....49..993E. doi:10.1130/G48549.1. S2CID236585231.
^Hilton, E. J.; Kovalchuk, O.; Podoplelova, N. (2021). "Sturgeons (Acipenseridae) from the Late Miocene of Ukraine, with a discussion of materials associated with Widhalm's (1886) nomen nudum, †Acipenser euhuso". Zootaxa. 5057 (3): 85–101. doi:10.11646/zootaxa.5057.3.4. PMID34811203. S2CID239555590.
^Meunier, F. J.; Alvarado-Ortega, J.; Machado, L. P. C.; Brito, P. M. (2021). "The paleohistology of bone and teeth in Cretaceous Pycnodontidae (Neopterygii: Pycnodontiformes): the case of Neoproscinetes penalvai and Tepexichthys aranguthyorum". Cybium. 45 (3): 193–203. doi:10.26028/cybium/2021-453-003.
^Arratia, G.; Schultze, H.-P.; Gouiric-Cavalli, S.; Quezada-Romegialli, C. (2021). "The intriguing † Atacamichthys fish from the Middle Jurassic of Chile – an amiiform or a teleosteomorph?". In Pradel, A.; Denton, J. S. S.; Janvier, P. (eds.). Ancient Fishes and their Living Relatives: a Tribute to John G. Maisey. Munich, Germany: Verlag Dr. Friedrich Pfeil. pp. 19–36. ISBN978-3-89937-269-4.
^Downs, J. P.; Barbosa, J.; Daeschler, E. B. (2021). "A new species of Eusthenodon (Sarcopterygii, Tristichopteridae) from the Upper Devonian (Famennian) of Pennsylvania, U.S.A., and a review of Eusthenodon taxonomy". Journal of Vertebrate Paleontology. 41 (3): e1976197. Bibcode:2021JVPal..41E6197D. doi:10.1080/02724634.2021.1976197. S2CID240453731.
^Renesto, S.; Magnani, F.; Stockar, R. (2021). "A new coelacanth specimen with elongate ribs from the Middle Triassic (Ladinian) Kalkschieferzone of Monte San Giorgio (Canton Ticino, Switzerland)". Rivista Italiana di Paleontologia e Stratigrafia. 127 (3): 689–700. doi:10.13130/2039-4942/16731.
^Hartung, J.; Sander, P. M.; Friedman, M.; Wintrich, T. (2021). "First record of mawsoniid coelacanths (Actinistia, Sarcopterygii) from the marine Rhaetian (Upper Triassic) of Bonenburg, Germany". Journal of Vertebrate Paleontology. 41 (2): e1931258. Bibcode:2021JVPal..41E1258H. doi:10.1080/02724634.2021.1931258. S2CID237517969.
^Ballen, G. A.; Jaramillo, C.; Dagosta, F. C. P.; de Pinna, M. C. C. (2021). "A fossil fish assemblage from the middle Miocene of the Cocinetas Basin, northern Colombia". Papers in Palaeontology. 8. doi:10.1002/spp2.1405. S2CID244439921.
