List of crocidurines
Species in mammal subfamily Crocidurinae
Greater white-toothed shrew (Crocidura russula )
Crocidurinae is a subfamily of small mammals in the shrew family Soricidae , which in turn is part of the order Eulipotyphla . A member of this family is called a crocidurine, or a white-toothed shrew. Crocidurinae is one of three subfamilies in Soricidae, along with the African shrews of Myosoricinae and the red-toothed shrews of Soricinae . They are found in Africa, Europe, and Asia, primarily in forests, savannas , shrublands , and grasslands , though some species can also be found in wetlands, rocky areas, or coastal areas. They range in size from the Etruscan shrew , one of the smallest mammal species at 3 cm (1 in) plus a 2 cm (1 in) tail,[ 1] to the hero shrew and Thor's hero shrew , at 15 cm (6 in) plus a 10 cm (4 in) tail. Crocidurines primarily eat invertebrates , small mammals, frogs, toads, and lizards. Almost no crocidurines have population estimates, but 21 species are categorized as endangered species , while the Andaman shrew , Christmas Island shrew , Harenna shrew , Jenkins's shrew , Nicobar shrew , and Wimmer's shrew are categorized as critically endangered .
The 235 extant species of Crocidurinae are divided into nine genera ; 192 of them are in Crocidura , the most species of any mammal genus.[ 2] A few extinct prehistoric Crocidurinae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[ 3]
Conventions
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species . Range maps are provided wherever possible; if a range map is not available, a description of the crocidurine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.
Classification
The subfamily Crocidurinae consists of nine genera: Crocidura , containing 192 species; Diplomesodon and Feroculus with a single species each; Paracrocidura , with three species; Ruwenzorisorex , with one species; Scutisorex , containing two species; Solisorex , with one species, Suncus , containing nineteen species; and Sylvisorex , containing fifteen species.
Subfamily Crocidurinae
Genus Crocidura (white-toothed shrews): 191 species
Genus Diplomesodon (piebald shrew): one species
Genus Feroculus (Kelaart's long-clawed shrew): one species
Genus Paracrocidura (large-headed shrews): three species
Genus Ruwenzorisorex (Ruwenzori shrew): one species
Genus Scutisorex (hero shrews): two species
Genus Solisorex (Pearson's long-clawed shrew): one species
Genus Suncus (musk shrews and pygmy shrews): nineteen species
Genus Sylvisorex (forest shrews): fifteen species
Crocidurines
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis , as supported by both the IUCN and the American Society of Mammalogists .[ 5]
Genus Crocidura โ Wagler , 1832 โ 192 species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
African black shrew
C. nigrofusca Matschie , 1895
Central and eastern Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 7] Diet : Invertebrates , small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 7]
African dusky shrew
C. caliginea Hollister , 1916
Central Africa
Size : 6โ8 cm (2โ3 in) long, plus 4โ6 cm (2 in) tail[ 8] Habitat : Forest[ 9] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 9]
African giant shrew
C. olivieri (Lesson , 1827)
C. o. anchietae
C. o. bueae
C. o. cara
C. o. cinereoaenea
C. o. darfurea
C. o. giffardi
C. o. guineensis
C. o. hansruppi
C. o. hedenborgiana
C. o. kivu
C. o. manni
C. o. martiensseni
C. o. nyansae
C. o. occidentalis
C. o. odorata
C. o. olivieri
C. o. spurelli
C. o. sururae
C. o. toritensis
C. o. zuleika
Sub-Saharan Africa
Size : 11โ14 cm (4โ6 in) long, plus 8โ10 cm (3โ4 in) tail[ 10] Habitat : Forest, savanna , and shrubland [ 11] Diet : Invertebrates[ 10]
LC
Unknown [ 11]
Andaman shrew
C. andamanensis Miller , 1902
South Andaman Island in India
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 12] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 12]
Andaman spiny shrew
C. hispida Thomas , 1913
South Andaman Island in India
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 13] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 13]
Annamite shrew
C. annamitensis Jenkins , Lunde , & Moncrieff , 2009
Vietnam
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 14] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 14]
Ansell's shrew
C. ansellorum Hutterer & Dippenaar , 1987
South-central Africa
Size : 5โ6 cm (2โ2 in) long, plus 4โ6 cm (2 in) tail[ 15] Habitat : Forest[ 16] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 16]
Arabian shrew
C. arabica Hutterer , 1988
Oman and Yemen
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 17] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 17]
Armenian shrew
C. armenica Gureev , 1963
Armenia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Rocky areas[ 18] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 18]
Asian gray shrew
C. attenuata H. Milne-Edwards , 1872
Eastern and southeastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest, shrubland, and grassland [ 19] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 19]
Asian lesser white-toothed shrew
C. shantungensis Miller , 1901
C. s. quelpartis
C. s. shantungensis
Eastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland and grassland[ 20] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 20]
Bailey's shrew
C. baileyi Osgood , 1936
Ethiopia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Grassland[ 21] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 21]
Bale shrew
C. bottegoides Hutterer & Yalden , 1990
Ethiopia
Size : 4โ6 cm (2 in) long, plus 4โ5 cm (2 in) tail[ 22] Habitat : Grassland and forest[ 23] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 23]
Banka shrew
C. vosmaeri Jentink , 1888
Bangka Island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 24] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 24]
Batak shrew
C. batakorum Hutterer , 2007
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 25] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 25]
Bates's shrew
C. batesi (Dollman , 1915)
Western Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 26] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 26]
Beccari's shrew
C. beccarii Dobson , 1886
Sumatra
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 27] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 27]
Beletta shrew
C. yaldeni Lavrenchenko , Voyta , & Hutterer , 2016
Ethiopia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 28] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 28]
Bicolored musk shrew
C. fuscomurina (Heuglin , 1865)
Sub-Saharan Africa
Size : 4โ8 cm (2โ3 in) long, plus 2โ5 cm (1โ2 in) tail[ 29] Habitat : Savanna, grassland, and desert[ 30] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 30]
Bicolored shrew
C. leucodon (Hermann , 1780)
Europe and western Asia
Size : 6โ9 cm (2โ4 in) long, plus 2โ5 cm (1โ2 in) tail[ 31] Habitat : Shrubland and grassland[ 32] Diet : Small mammals, frogs, toads, lizards, and invertebrates[ 31]
LC
Unknown [ 32]
Black-footed shrew
C. nigripes Miller & Hollister , 1921
C. n. lipara
C. n. nigripes
Sulawesi island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Blackish white-toothed shrew
C. nigricans Bocage , 1889
Angola
Size : 6โ9 cm (2โ4 in) long, plus 4โ7 cm (2โ3 in) tail[ 34] Habitat : Unknown[ 35] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 35]
Bornean shrew
C. foetida Peters , 1870
C. f. doriae
C. f. foetida
C. f. kelabit
Borneo
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 36] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 36]
Bottego's shrew
C. bottegi Thomas , 1898
Ethiopia
Size : 4โ6 cm (2 in) long, plus 2โ3 cm (1 in) tail[ 37] Habitat : Unknown[ 38] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 38]
Buettikofer's shrew
C. buettikoferi Jentink , 1888
West-central Africa
Size : 7โ10 cm (3โ4 in) long, plus 5โ7 cm (2โ3 in) tail[ 39] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Butiaba naked-tailed shrew
C. littoralis Heller , 1910
Central Africa
Size : 8โ10 cm (3โ4 in) long, plus 6โ8 cm (2โ3 in) tail[ 41] Habitat : Forest[ 42] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 42]
Cameroonian shrew
C. picea Sanderson , 1940
West-central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and grassland[ 43] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 43]
Canarian shrew
C. canariensis Hutterer , Lรณpez-Jurado , & Vogel , 1987
Canary Islands
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Coastal marine, desert, and shrubland[ 44] Diet : Snails and insects[ 44]
EN
Unknown [ 44]
Caspian shrew
C. caspica Thomas , 1907
Iran and Azerbaijan
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 45] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 45]
Chinese white-toothed shrew
C. rapax Allen , 1923
C. r. kurodai
C. r. lutaoensis
C. r. rapax
C. r. tadae
Eastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 46] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 46]
Christmas Island shrew
C. trichura Dobson , 1889
Christmas Island
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 47] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
0โ200 [ 47]
Cinderella shrew
C. cinderella Thomas , 1911
Western Africa
Size : 5โ7 cm (2โ3 in) long, plus 4โ5 cm (2โ2 in) tail[ 48] Habitat : Savanna[ 49] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 49]
Congo white-toothed shrew
C. congobelgica Hollister , 1916
Central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 50] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 50]
Cranbrook's white-toothed shrew
C. cranbrooki Jenkins , Lunde , & Moncrieff , 2009
Northern Myanmar
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 51] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 51]
Cretan shrew
C. zimmermanni Wettstein , 1953
Island of Crete in Greece
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown and rocky areas[ 52] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 52]
Crosse's shrew
C. crossei Thomas , 1895
Western Africa
Size : 6โ7 cm (2โ3 in) long, plus 4โ6 cm (2 in) tail[ 53] Habitat : Forest[ 54] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 54]
Cyrenaica shrew
C. aleksandrisi Vesmanis , 1977
Northern Libya
Size : 5โ7 cm (2โ3 in) long, plus 3โ5 cm (1โ2 in) tail[ 55] Habitat : Shrubland[ 56] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 56]
Dent's shrew
C. denti Dollman , 1915
Western and central Africa
Size : About 6 cm (2 in) long, plus 5 cm (2 in) tail[ 57] Habitat : Forest and savanna[ 58] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 58]
Desert musk shrew
C. smithii Thomas , 1895
C. s. debalsaci
C. s. smithii
Western and Eastern Africa
Size : 6โ9 cm (2โ4 in) long, plus 3โ6 cm (1โ2 in) tail[ 59] Habitat : Savanna[ 60] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 60]
Desperate shrew
C. desperata Hutterer , Jenkins , & Verheyen , 1991
Tanzania
Size : 8โ11 cm (3โ4 in) long, plus about 7 cm (3 in) tail[ 61] Habitat : Forest[ 62] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 62]
Dhofar shrew
C. dhofarensis (Hutterer , 1988)
Yemen and Oman
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 63] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 63]
Doucet's musk shrew
C. douceti Heim de Balsac , 1958
Western Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 64] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 64]
Dsinezumi shrew
C. dsinezumi (Temminck , 1842)
Japan
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland, grassland, and inland wetlands[ 65] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 65]
East African highland shrew
C. allex Osgood , 1910
Kenya and Tanzania
Size : 5โ7 cm (2โ3 in) long, plus 4โ6 cm (2 in) tail[ 66] Habitat : Grassland and inland wetlands[ 67] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 67]
Egyptian pygmy shrew
C. religiosa (Geoffroy , 1827)
Northern Egypt
Size : 4โ7 cm (2โ3 in) long, plus 2โ4 cm (1โ2 in) tail[ 68] Habitat : [ 69] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 69]
Eisentraut's shrew
C. eisentrauti Heim de Balsac , 1957
Cameroon
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Grassland and forest[ 70] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 70]
Elgon shrew
C. elgonius Osgood , 1910
Kenya and Tanzania
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 71] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 71]
Elongated shrew
C. elongata Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and shrubland[ 72] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 72]
Fingui white-toothed shrew
C. fingui Cerรญaco et al., 2015
Prรญncipe in Sรฃo Tomรฉ and Prรญncipe
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 73] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 73]
Fischer's shrew
C. fischeri Pagenstecher , 1885
Kenya and Tanzania
Size : 6โ10 cm (2โ4 in) long, plus 4โ6 cm (2 in) tail[ 74] Habitat : SavannaDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Flat-headed shrew
C. planiceps Heller , 1910
West-central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Inland wetlands[ 76] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 76]
Flower's shrew
C. floweri Dollman , 1915
Northern Egypt
Size : 5โ8 cm (2โ3 in) long, plus 5โ6 cm (2โ2 in) tail[ 77] Habitat : Artificial terrainDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Fox's shrew
C. foxi Dollman , 1915
Western and central Africa
Size : 9โ12 cm (4โ5 in) long, plus 5โ8 cm (2โ3 in) tail[ 79] Habitat : Savanna and forest[ 80] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 80]
Fraser's musk shrew
C. poensis (Fraser , 1843)
Western Africa
Size : 7โ10 cm (3โ4 in) long, plus 4โ7 cm (2โ3 in) tail[ 81] Habitat : Shrubland and forest[ 82] Diet : Arthropods and other invertebrates[ 81]
LC
Unknown [ 82]
Gathorne's shrew
C. gathornei Jenkins , 2013
Himalayas in India
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland and grassland[ 83] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 83]
Glass's shrew
C. glassi Heim de Balsac , 1966
Ethiopia
Size : 6โ9 cm (2โ4 in) long, plus 4โ6 cm (2 in)tail[ 84] Habitat : Shrubland, grassland, and inland wetlands[ 85] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 85]
Gmelin's white-toothed shrew
C. gmelini Pallas , 1811
Central Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland and grassland[ 86] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 86]
Goliath shrew
C. goliath Thomas , 1906
Central Africa
Size : 15โ18 cm (6โ7 in) long, plus 10โ14 cm (4โ6 in) tail[ 87] Habitat : Forest[ 88] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 88]
Gracile naked-tailed shrew
C. maurisca Thomas , 1904
Central Africa
Size : 7โ10 cm (3โ4 in) long, plus 6โ8 cm (2โ3 in) tail[ 89] Habitat : Inland wetlands and forest[ 90] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 90]
Grasse's shrew
C. grassei Brosset , Dubost , & Heim de Balsac , 1965
West-central Africa
Size : 7โ9 cm (3โ4 in) long, plus 6โ8 cm (2โ3 in) tail[ 91] Habitat : Forest[ 92] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 92]
Greater Mindanao shrew
C. grandis Miller , 1911
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 93] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 93]
Greater red musk shrew
C. flavescens (Geoffroy , 1827)
Southern Africa
Size : 8โ12 cm (3โ5 in) long, plus 3โ6 cm (1โ2 in) tail[ 94] Habitat : Savanna and grassland[ 95] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 95]
Greater white-toothed shrew
C. russula (Hermann , 1780)
Europe and northwestern Africa
Size : 6โ9 cm (2โ4 in) long, plus 3โ5 cm (1โ2 in) tail[ 96] Habitat : Shrubland[ 97] Diet : Invertebrates, as well as small mammals and small lizards[ 96]
LC
Unknown [ 97]
Greenwood's shrew
C. greenwoodi Heim de Balsac , 1966
Somalia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest, savanna, and shrubland[ 98] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 98]
Guramba shrew
C. phaeura Osgood , 1936
Ethiopia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 99] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 99]
Guy's shrew
C. guy Jenkins , Lunde , & Moncrieff , 2009
Vietnam
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 100] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 100]
Gรผldenstรคdt's shrew
C. gueldenstaedtii (Pallas , 1811)
Europe and western Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland, grassland, inland wetlands, rocky areas, intertidal marine , and coastal marine[ 101] Diet : Insects, as well as molluscs, amphipods , annelids , and small vertebrates[ 101]
LC
Unknown [ 101]
Hainan Island shrew
C. wuchihensis Sung , 1966
Southern China and northern Vietnam
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 102] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 102]
Harenna shrew
C. harenna Hutterer & Yalden , 1990
Ethiopia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 103] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 103]
Heather shrew
C. erica Dollman , 1915
Angola
Size : About 10 cm (4 in) long, plus 5 cm (2 in) tail[ 104] Habitat : Unknown[ 105] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 105]
Hildegarde's shrew
C. hildegardeae Thomas , 1904
Central Africa
Size : 6โ8 cm (2โ3 in) long, plus 3โ6 cm (1โ2 in) tail[ 106] Habitat : Forest[ 107] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 107]
Hill's shrew
C. hilliana Jenkins & Smith , 1995
Thailand and Laos
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and shrubland[ 108] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 108]
Horsfield's shrew
C. horsfieldii (Tomes , 1856)
Scattered southern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 109] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 109]
Hun shrew
C. attila Dollman , 1915
Central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 110] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 110]
Hutan shrew
C. hutanis Ruedi & Vogel , 1995
Sumatra island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 111] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 111]
Indochinese shrew
C. indochinensis Kloss , 1922
Southeastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 112] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 112]
Iranian shrew
C. susiana Redding & Lay , 1978
Iran
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 113] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 113]
Jackass shrew
C. arispa Spitzenberger , 1971
Turkey
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Rocky areas[ 114] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 114]
Jackson's shrew
C. jacksoni Thomas , 1904
East-central Africa
Size : 5โ9 cm (2โ4 in) long, plus 4โ7 cm (2โ3 in) tail[ 115] Habitat : Forest[ 116] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 116]
Javan ghost shrew
C. umbra Demos , Achmadi , Handika , Maharadatunkamsi , Rowe , & Esselstyn , 2016
Java island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 117] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 117]
Javanese shrew
C. maxi Sody , 1936
Indonesia and East Timor
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 118] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 118]
Jenkins's shrew
C. jenkinsi Chakraborty , 1978
South Andaman Island in India
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 119] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 119]
Jouvenet's shrew
C. jouvenetae Heim de Balsac , 1958
Western Africa
Size : 6โ9 cm (2โ4 in) long, plus 4โ7 cm (2โ3 in) tail[ 120] Habitat : Forest[ 121] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 121]
Kashmir white-toothed shrew
C. pullata Miller , 1911
Northern India and Pakistan
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 122] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 122]
Katinka's shrew
C. katinka Bate , 1937
Western Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 123] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 123]
Kilimanjaro shrew
C. monax Thomas , 1910
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 124] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 124]
Kinabalu shrew
C. baluensis Thomas , 1898
Northern Borneo
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest, shrubland, and grassland[ 125] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 125]
Kivu long-haired shrew
C. lanosa Heim de Balsac , 1968
Central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and inland wetlands[ 126] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 126]
Kivu shrew
C. kivuana Heim de Balsac , 1968
Central Africa
Size : 6โ9 cm (2โ4 in) long, plus 6โ8 cm (2โ3 in) tail[ 127] Habitat : Forest and inland wetlands[ 128] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 128]
Lamotte's shrew
C. lamottei Heim de Balsac , 1968
Western Africa
Size : 8โ9 cm (3โ4 in) long, plus 3โ6 cm (1โ2 in) tail[ 127] Habitat : Forest and savanna[ 129] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 129]
Large-headed forest shrew
C. grandiceps Hutterer , 1983
Western Africa
Size : 9โ12 cm (4โ5 in) long, plus 5โ8 cm (2โ3 in) tail[ 130] Habitat : Forest[ 131] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 131]
Latona's shrew
C. latona Hollister , 1916
Democratic Republic of the Congo
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 132] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 132]
Lesser Ryukyu shrew
C. watasei Kuroda , 1924
Southern Japan
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland, grassland, and inland wetlands[ 133] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 133]
Lesser gray-brown musk shrew
C. silacea Thomas , 1895
Southern Africa
Size : 6โ8 cm (2โ3 in) long, plus 4โ6 cm (2 in) tail[ 134] Habitat : Rocky areas, grassland, shrubland, and savanna[ 135] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 135]
Lesser red musk shrew
C. hirta Peters , 1852
Southern and southeastern Africa
Size : 6โ10 cm (2โ4 in) long, plus 3โ6 cm (1โ2 in) tail[ 136] Habitat : Savanna, grassland, and desert[ 137] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 137]
Lesser rock shrew
C. serezkyensis Laptev , 1929
Western Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Unknown[ 138] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 138]
Lesser white-toothed shrew
C. suaveolens (Pallas , 1811)
Europe and Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest, shrubland, grassland, rocky areas, and coastal/supratidal marine [ 139] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 139]
Long-footed shrew
C. crenata Brosset , Dubost , & Heim de Balsac , 1965
West-central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Long-tailed musk shrew
C. dolichura Peters , 1876
Central Africa
Size : 6โ7 cm (2โ3 in) long, plus 7โ8 cm (3 in) tail[ 141] Habitat : Forest[ 142] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 142]
Lucina's shrew
C. lucina Dippenaar , 1980
Ethiopia
Size : 8โ10 cm (3โ4 in) long, plus 4โ6 cm (2 in) tail[ 143] Habitat : Grassland and inland wetlands[ 144] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 144]
Ludia's shrew
C. ludia Hollister , 1916
Central Africa
Size : 6โ7 cm (2โ3 in) long, plus 5โ6 cm (2โ2 in) tail[ 145] Habitat : Forest[ 146] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 146]
Luzon shrew
C. grayi Dobson , 1890
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 147] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 147]
Lwiro shrew
C. lwiroensis Peterhans & Hutterer , 2013
Democratic Republic of the Congo
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 148] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 148]
MacArthur's shrew
C. macarthuri St. Leger , 1934
Eastern Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : SavannaDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
MacMillan's shrew
C. macmillani Dollman , 1915
Ethiopia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and savanna[ 150] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 150]
Makwassie musk shrew
C. maquassiensis Roberts , 1946
Southern Africa
Size : 5โ8 cm (2โ3 in) long, plus 4โ5 cm (2โ2 in) tail[ 151] Habitat : Grassland and rocky areas[ 152] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 152]
Malayan shrew
C. malayana Robinson & Kloss , 1911
Malaysia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 153] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 153]
Mamfe shrew
C. virgata Sanderson , 1940
West-central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and grassland[ 154] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 154]
Manenguba shrew
C. manengubae Hutterer , 1982
West-central Africa
Size : 7โ9 cm (3โ4 in) long, plus 6โ7 cm (2โ3 in) tail[ 151] Habitat : Forest, shrubland, and grassland[ 155] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 155]
Mauritanian shrew
C. lusitania Dollman , 1915
Western and eastern Africa
Size : 4โ7 cm (2โ3 in) long, plus 2โ4 cm (1โ2 in) tail[ 156] Habitat : Savanna and shrubland[ 157] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 157]
Mduma's shrew
C. mdumai Stanley , Hutterer , Giarla , & Esselstyn , 2015
Tanzania
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 158] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 158]
Mindanao shrew
C. beatus Miller , 1910
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 159] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 159]
Mindoro shrew
C. mindorus Miller , 1910
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 160] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 160]
Montane white-toothed shrew
C. montis Thomas , 1906
Eastern Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest, shrubland, and grassland[ 161] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 161]
Moonshine shrew
C. luna Dollman , 1910
Southeastern Africa
Size : 6โ11 cm (2โ4 in) long, plus 3โ7 cm (1โ3 in) tail[ 156] Habitat : Forest[ 162] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 162]
Mossy forest shrew
C. musseri Ruedi & Vogel , 1995
Sulawesi island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 163] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 163]
Mount Nimba giant forest shrew
C. nimbasilvanus Hutterer , 2003
Western Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 164] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 164]
Munissi's shrew
C. munissii Stanley , Hutterer , Giarla , & Esselstyn , 2015
Tanzania
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 165] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 165]
Narrow-headed shrew
C. stenocephala Heim de Balsac & Dieterlen , 1979
Central Africa
Size : 8โ11 cm (3โ4 in) long, plus 6โ9 cm (2โ4 in) tail[ 166] Habitat : Inland wetlands[ 167] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 167]
Negev shrew
C. ramona Ivanitskaya , Shenbrot , & Nevo , 1996
Israel
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Desert[ 168] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 168]
Negros shrew
C. negrina Rabor , 1952
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 169] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 169]
Nicobar shrew
C. nicobarica Miller , 1902
Great Nicobar Island in India
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 170] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown [ 170]
Nigerian shrew
C. nigeriae Dollman , 1915
West-central Africa
Size : 9โ11 cm (4 in) long, plus 5โ7 cm (2โ3 in) tail[ 171] Habitat : Forest[ 172] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 172]
Nimba shrew
C. nimbae Heim de Balsac , 1956
Western Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 173] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 173]
Niobe's shrew
C. niobe Thomas , 1906
Central Africa
Size : 6โ9 cm (2โ4 in) long, plus 5โ7 cm (2โ3 in) tail[ 174] Habitat : Forest and grassland[ 175] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 175]
North African white-toothed shrew
C. pachyura Kรผster , 1835
Northern Africa and southern Europe
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland[ 176] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 176]
Nyiro shrew
C. macowi Dollman , 1915
Kenya
Size : 6โ8 cm (2โ3 in) long, plus 5โ6 cm (2โ2 in) tail[ 177] Habitat : Unknown[ 178] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 178]
Oriental shrew
C. orientalis Jentink , 1890
C. o. lawuana
C. o. orientalis
Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 179] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 179]
Palawan shrew
C. palawanensis Taylor , 1934
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and shrubland[ 180] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 180]
Pale gray shrew
C. pergrisea Miller , 1913
Pakistan
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 181] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 181]
Panay shrew
C. panayensis Hutterer , 2007
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 182] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 182]
Peninsular shrew
C. negligens Kloss , 1914
Southeastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 183] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 183]
Peters's musk shrew
C. gracilipes Peters , 1870
Tanzania
Size : About 6 cm (2 in) long, plus 5 cm (2 in) tail[ 87] Habitat : UnknownDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Phan Luong's shrew
C. phanluongi Jenkins , Abramov , Rozhnov , & Olsson , 2010
Vietnam and Cambodia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 185] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 185]
Phu Hoc shrew
C. phuquocensis Abramov , Jenkins , Rozhnov , & Kalinin , 2008
Phรบ Quแปc island in Vietnam
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 186] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 186]
Pitman's shrew
C. pitmani Barclay, 1932
Zambia
Size : About 5 cm (2 in) long, plus 4 cm (2 in) tail[ 187] Habitat : Unknown[ 188] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 188]
Polia's shrew
C. polia Hollister , 1916
Democratic Republic of the Congo
Size : About 6 cm (2 in) long, plus 7 cm (3 in) tail[ 81] Habitat : Unknown[ 189] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 189]
Rainey's shrew
C. raineyi Heller , 1912
Kenya
Size : 9โ10 cm (4 in) long, plus 5โ7 cm (2โ3 in) tail[ 190] Habitat : Forest[ 191] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 191]
Reddish-gray musk shrew
C. cyanea (Duvernoy , 1838)
Southern Africa
Size : 6โ10 cm (2โ4 in) long, plus 3โ7 cm (1โ3 in) tail[ 192] Habitat : Forest, grassland, and caves[ 193] Diet : Invertebrates[ 193]
LC
Unknown [ 193]
Roosevelt's shrew
C. roosevelti (Heller , 1910)
Central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Savanna[ 194] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 194]
Ryukyu shrew
C. orii Kuroda , 1924
Southern Japan
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 195] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 195]
Sa Pa shrew
C. sapaensis Jenkins , Abramov , Bannikova , & Rozhnov , 2013
Vietnam
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and grassland[ 196] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 196]
Saharan shrew
C. tarfayensis Vesmanis & Vesmanis , 1980
Mauritania and Morocco
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Rocky areas and desert[ 197] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 197]
Sahelian tiny shrew
C. pasha Dollman , 1915
Scattered northern Africa
Size : 4โ6 cm (2 in) long, plus 2โ4 cm (1โ2 in) tail[ 198] Habitat : Savanna[ 199] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 199]
Savanna dwarf shrew
C. nanilla Thomas , 1909
Western and eastern Africa
Size : 4โ6 cm (2โ2 in) long, plus 3โ5 cm (1โ2 in) tail[ 200] Habitat : Savanna[ 201] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 201]
Savanna path shrew
C. viaria (Geoffroy , 1834)
Western, central, and eastern Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and savanna[ 202] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 202]
Savanna shrew
C. fulvastra (Sundevall , 1843)
Central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Savanna[ 203] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 203]
Savanna swamp shrew
C. longipes Hutterer & Happold , 1983
West-central Africa
Size : 9โ12 cm (4โ5 in) long, plus 5โ7 cm (2โ3 in) tail[ 204] Habitat : Inland wetlands[ 205] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 205]
Siberian shrew
C. sibirica Dukelsky , 1930
Central Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest, grassland, inland wetlands, and desert[ 206] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 206]
Sibuyan shrew
C. ninoyi Esselstyn & Goodman , 2010
Philippines
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 207] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 207]
Sicilian shrew
C. sicula Miller , 1900
C. s. aegatensis
C. s. calypso
C. s. esuae
C. s. sicula
Italy and Malta
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Shrubland[ 208] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 208]
Sinharaja white-toothed shrew
C. hikmiya Meegaskumbura , Meegaskumbura , Pethiyagoda , Manamendra-Arachchi , & Schneider , 2007
Sri Lanka
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 209] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 209]
Small-footed shrew
C. parvipes Osgood , 1910
Central Africa
Size : 6โ9 cm (2โ4 in) long, plus 3โ6 cm (1โ2 in) tail[ 210] Habitat : Forest, savanna, and shrublandDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Smoky white-toothed shrew
C. fumosa Thomas , 1904
Kenya
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 212] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 212]
Sokolov's shrew
C. sokolovi Jenkins , Abramov , Rozhnov , & Makarova , 2007
Vietnam
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 213] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 213]
Somali dwarf shrew
C. nana Dobson , 1890
Ethiopia and Somalia
Size : About 4 cm (2 in) long, plus 3 cm (1 in) tail[ 214] Habitat : Grassland[ 215] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 215]
Somali shrew
C. somalica Thomas , 1895
Eastern Africa
Size : 6โ8 cm (2โ3 in) long, plus 4โ5 cm (2โ2 in) tail[ 166] Habitat : Savanna[ 216] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 216]
Southeast Asian shrew
C. fuliginosa (Blyth , 1856)
Southeastern Asian
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest, shrubland, grassland, and rocky areas[ 217] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 217]
Sri Lankan long-tailed shrew
C. miya Phillips , 1929
Sri Lanka
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 218] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 218]
Sulawesi shrew
C. lea Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 219] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 219]
Sulawesi tiny shrew
C. levicula Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 220] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 220]
Sulawesi white-handed shrew
C. rhoditis Miller & Hollister , 1921
Sulawesi island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 221] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 221]
Sumatran giant shrew
C. lepidura Lyon , 1908
Sumatra island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and shrubland[ 222] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 222]
Sumatran long-tailed shrew
C. paradoxura Dobson , 1886
Sumatra island in Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 223] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 223]
Sunda shrew
C. monticola Peters , 1870
Southeastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 224] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 224]
Swamp musk shrew
C. mariquensis (Smith , 1844)
C. m. mariquensis
C. m. neavei
C. m. shortridgei
Southern Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Inland wetlands[ 225] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 225]
Sรฃo Tomรฉ shrew
C. thomensis (Bocage , 1887)
Sรฃo Tomรฉ Island in Sรฃo Tomรฉ and Prรญncipe
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 226] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 226]
Taiwanese gray shrew
C. tanakae Kuroda , 1938
Eastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest and grassland[ 227] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 227]
Tanzanian shrew
C. tansaniana Hutterer , 1986
Tanzania
Size : 8โ11 cm (3โ4 in) long, plus 5โ8 cm (2โ3 in) tail[ 228] Habitat : Forest[ 229] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
NT
Unknown [ 229]
Tarella shrew
C. tarella Dollman , 1915
Central Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 230] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
EN
Unknown [ 230]
Telford's shrew
C. telfordi Hutterer , 1986
Tanzania
Size : 10โ11 cm (4 in) long, plus 7โ9 cm (3โ4 in) tail[ 231] Habitat : Forest[ 232] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 232]
Thalia's shrew
C. thalia Dippenaar , 1980
Ethiopia
Size : 7โ10 cm (3โ4 in) long, plus 5โ7 cm (2โ3 in) tail[ 231] Habitat : Forest, savanna, shrubland, grassland, and inland wetlands[ 233] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 233]
Therese's shrew
C. theresae Heim de Balsac , 1968
Western Africa
Size : 7โ11 cm (3โ4 in) long, plus 3โ6 cm (1โ2 in) tail[ 234] Habitat : Forest and savanna[ 235] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 235]
Thick-tailed shrew
C. brunnea Jentink , 1888
C. b. brunnea
C. b. pudjonica
Indonesia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 236] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 236]
Timor shrew
C. tenuis (Mรผller , 1840)
Timor
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 237] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 237]
Turbo shrew
C. turba Dollman , 1910
Central Africa
Size : 9โ11 cm (4 in) long, plus 5โ6 cm (2โ2 in) tail[ 238] Habitat : Forest[ 239] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 239]
Ugandan lowland shrew
C. selina (Dollman , 1915)
Kenya and Uganda
Size : 7โ9 cm (3โ4 in) long, plus 5โ7 cm (2โ3 in) tail[ 240] Habitat : Forest and inland wetlands[ 241] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 241]
Ugandan musk shrew
C. mutesae Heller , 1910
Central Africa
Size : About 11 cm (4 in) long, plus 6 cm (2 in) tail[ 214] Habitat : Forest[ 242] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 242]
Ultimate shrew
C. ultima Dollman , 1915
Kenya
Size : About 9 cm (4 in) long, plus 6 cm (2 in) tail[ 243] Habitat : Forest[ 244] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 244]
Upemba shrew
C. zimmeri Osgood , 1936
Democratic Republic of the Congo
Size : 11โ13 cm (4โ5 in) long, plus 4โ6 cm (2โ2 in) tail[ 245] Habitat : Unknown[ 246] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown [ 246]
Usambara shrew
C. usambarae Dippenaar , 1980
Tanzania
Size : 7โ9 cm (3โ4 in) long, plus 5โ7 cm (2โ3 in) tail[ 247] Habitat : Forest[ 248] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
VU
Unknown [ 248]
Ussuri white-toothed shrew
C. lasiura (Dobson , 1890)
Eastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Grassland[ 249] Diet : Insects and other invertebrates, as well as small vertebrates[ 249]
LC
Unknown [ 249]
Voi shrew
C. voi Osgood , 1910
Central and eastern Africa
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Savanna[ 250] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 250]
Voracious shrew
C. vorax Allen , 1923
Southeastern Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Forest[ 251] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 251]
West African long-tailed shrew
C. muricauda (Miller , 1900)
Western Africa
Size : 5โ7 cm (2โ3 in) long, plus tail[ 252] Habitat : Forest[ 253] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 253]
West African pygmy shrew
C. obscurior Heim de Balsac , 1958
Western Africa
Size : 4โ5 cm (2โ2 in) long, plus 3โ4 cm (1โ2 in) tail[ 254] Habitat : Forest[ 255] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 255]
Whitaker's shrew
C. whitakeri De Winton , 1898
Northwestern Africa
Size : 5โ7 cm (2โ3 in) long, plus 2โ4 cm (1โ2 in) tail[ 256] Habitat : Coastal marine, intertidal marine, and shrublandDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown
Wimmer's shrew
C. wimmeri Heim de Balsac & Aellen , 1958
Ivory Coast
Size : 8โ9 cm (3โ4 in) long, plus 5โ6 cm (2โ2 in) tail[ 258] Habitat : ForestDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
CR
Unknown
Xanthippe's shrew
C. xantippe Osgood , 1910
Kenya and Tanzania
Size : 8โ10 cm (3โ4 in) long, plus 5โ7 cm (2โ3 in) tail[ 260] Habitat : Savanna and shrubland[ 261] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 261]
Yankari shrew
C. yankariensis Hutterer & Jenkins , 1980
Central and eastern Africa
Size : 5โ7 cm (2โ3 in) long, plus 3โ5 cm (1โ2 in) tail[ 260] Habitat : Savanna[ 262] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 262]
Zaphir's shrew
C. zaphiri Dollman , 1915
Ethiopia and Kenya
Size : About 10 cm (4 in) long, plus 6 cm (2 in) tail[ 263] Habitat : UnknownDiet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
DD
Unknown
Zarudny's rock shrew
C. zarudnyi Ognew , 1928
Central Asia
Size : 4โ18 cm (2โ7 in) long, plus 4โ11 cm (2โ4 in) tail[ 6] Habitat : Desert[ 265] Diet : Invertebrates, small mammals, frogs, toads, and lizards[ 6]
LC
Unknown [ 265]
Genus Diplomesodon โ Brandt , 1852 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Piebald shrew
D. pulchellum (Lichtenstein , 1823)
Central Asia
Size : 5โ8 cm (2โ3 in) long, plus 2โ4 cm (1โ2 in) tail[ 266] Habitat : Desert[ 267] Diet : Insects, as well as small lizards[ 267]
LC
Unknown [ 267]
Genus Feroculus โ Kelaart , 1852 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Kelaart's long-clawed shrew
F. feroculus (Kelaart , 1850)
Southern India and Sri Lanka
Size : 10โ12 cm (4โ5 in) long, plus 5โ8 cm (2โ3 in) tail[ 268] Habitat : Forest, grassland, and inland wetlands[ 269] Diet : Earthworms, small vertebrates, and plants[ 268]
EN
Unknown [ 269]
Genus Ruwenzorisorex โ Hutterer , 1986 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Ruwenzori shrew
R. suncoides (Osgood , 1936)
Central Africa
Size : 9โ10 cm (4 in) long, plus 5โ7 cm (2โ3 in) tail[ 274] Habitat : Inland wetlands and forest[ 275] Diet : Mollusks and small vertebrates[ 268]
VU
Unknown [ 275]
Genus Scutisorex โ Thomas , 1910 โ two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Hero shrew
S. somereni (Thomas , 1910)
Central Africa
Size : 12โ15 cm (5โ6 in) long, plus 6โ10 cm (2โ4 in) tail[ 276] Habitat : Forest[ 277] Diet : Insects, earthworms, and small vertebrates[ 276]
LC
Unknown [ 277]
Thor's hero shrew
S. thori Stanley , Malekani , &Gambalemoke , 2013
Democratic Republic of the Congo
Size : 12โ15 cm (5โ6 in) long, plus 6โ10 cm (2โ4 in) tail[ 278] Habitat : Forest[ 279] Diet : Small animals and plants[ 278]
DD
Unknown [ 279]
Genus Solisorex โ Thomas , 1924 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Pearson's long-clawed shrew
S. pearsoni Thomas , 1924
Sri Lanka
Size : 12โ15 cm (5โ6 in) long, plus 5โ7 cm (2โ3 in) tail[ 266] Habitat : Forest and grassland[ 280] Diet : Insects, earthworms, and small vertebrates[ 266]
EN
Unknown [ 280]
Genus Suncus โ Ehrenberg , 1832 โ eighteen species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Anderson's shrew
S. stoliczkanus (Anderson , 1877)
Southern Asia
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest and grassland[ 282] Diet : Insects[ 281]
LC
Unknown [ 282]
Asian highland shrew
S. montanus (Kelaart , 1850)
Southern India and Sri Lanka
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest and grassland[ 283] Diet : Insects[ 281]
LC
Unknown [ 283]
Asian house shrew
S. murinus (Linnaeus , 1766)
Asia and Africa (introduced in red)
Size : 10โ5 cm (4โ2 in) long, plus 5โ9 cm (2โ4 in) tail[ 284] Habitat : Forest, savanna, shrubland, grassland, and inland wetlands[ 285] Diet : Insects and small mammals, as well as plants[ 286]
LC
Unknown [ 285]
Black shrew
S. ater (Medway , 1965)
Northern Borneo
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest[ 287] Diet : Insects[ 281]
DD
Unknown [ 287]
Bornean pygmy shrew
S. hosei (Thomas , 1893)
Borneo
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest[ 288] Diet : Insects[ 281]
DD
Unknown [ 288]
Climbing shrew
S. megalura (Jentink , 1888)
Sub-Saharan Africa
Size : 5โ6 cm (2 in) long, plus 8โ10 cm (3โ4 in) tail[ 284] Habitat : Forest and savanna[ 289] Diet : Insects[ 284]
LC
Unknown [ 289]
Day's shrew
S. dayi (Dodson , 1888)
Southern India
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest and grassland[ 290] Diet : Insects[ 281]
EN
Unknown [ 290]
Etruscan shrew
S. etruscus (Savi , 1822)
Europe, northern Africa, and Asia
Size : 3โ5 cm (1โ2 in) long, plus 2โ3 cm (1 in) tail[ 1] Habitat : Forest, shrubland, grassland, and desert[ 291] Diet : Insects[ 292]
LC
Unknown [ 291]
Flores shrew
S. mertensi Kock , 1974
Flores island in Indonesia
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest[ 293] Diet : Insects[ 281]
EN
Unknown [ 293]
Greater dwarf shrew
S. lixus (Thomas , 1898)
Southern Africa
Size : 5โ9 cm (2โ4 in) long, plus 3โ6 cm (1โ2 in) tail[ 281] Habitat : Forest, savanna, shrubland, and grassland[ 294] Diet : Insects[ 281]
LC
Unknown [ 294]
Hutu-Tutsi dwarf shrew
S. hututsi Peterhans & Hutterer , 2009
Central Africa
Size : 4โ6 cm (2 in) long, plus 2โ3 cm (1 in) tail[ 281] Habitat : Forest and shrubland[ 295] Diet : Insects[ 281]
DD
Unknown [ 295]
Jungle shrew
S. zeylanicus Phillips , 1928
Sri Lanka
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest[ 296] Diet : Insects[ 281]
EN
Unknown [ 296]
Least dwarf shrew
S. infinitesimus (Heller , 1912)
South Africa and Kenya
Size : 5โ7 cm (2โ3 in) long, plus 2โ4 cm (1โ2 in) tail[ 281] Habitat : Forest and grassland[ 297] Diet : Insects[ 281]
LC
Unknown [ 297]
Lesser dwarf shrew
S. varilla (Thomas , 1895)
Southern Africa
Size : 5โ7 cm (2โ3 in) long, plus 2โ5 cm (1โ2 in) tail[ 281] Habitat : Forest, savanna, shrubland, and grassland[ 298] Diet : Insects[ 281]
LC
Unknown [ 298]
Malayan pygmy shrew
S. malayanus (Kloss , 1917)
Malaysia and Thailand
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest[ 299] Diet : Insects[ 281]
LC
Unknown [ 299]
Remy's pygmy shrew
S. remyi Brosset , Dubost , & Heim de Balsac , 1965
West-central Africa
Size : 4โ5 cm (2 in) long, plus 1โ3 cm (0.4โ1.2 in) tail[ 281] Habitat : Forest[ 300] Diet : Insects[ 281]
LC
Unknown [ 300]
Sri Lankan shrew
S. fellowesgordoni Phillips , 1932
Sri Lanka
Size : 5โ9 cm (2โ4 in) long, plus 3โ10 cm (1โ4 in) tail[ 281] Habitat : Forest and grassland[ 301] Diet : Insects[ 281]
EN
Unknown [ 301]
Taita shrew
S. aequatorius (Heller , 1912)
Kenya
Size : 6โ9 cm (2โ4 in) long, plus 5โ7 cm (2โ3 in) tail[ 302] Habitat : Forest[ 303] Diet : Insects[ 304]
EN
Unknown [ 303]
Genus Sylvisorex โ Thomas , 1904 โ fifteen species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Bamenda pygmy shrew
S. silvanorum (Hutterer , Riegert , & Sedlรกฤek , 2009)
Cameroon
Size : 4โ10 cm (2โ4 in) long, plus 4โ9 cm (2โ4 in) tail[ 274] Habitat : Forest and grassland[ 305] Diet : Insects[ 306]
DD
Unknown [ 305]
Bioko forest shrew
S. isabellae Heim de Balsac , 1968
West-central Africa
Size : 4โ10 cm (2โ4 in) long, plus 4โ9 cm (2โ4 in) tail[ 274] Habitat : Forest and grassland[ 307] Diet : Insects[ 306]
VU
Unknown [ 307]
Cameroonian forest shrew
S. camerunensis Heim de Balsac , 1968
West-central Africa
Size : 5โ6 cm (2 in) long, plus 5โ7 cm (2โ3 in) tail[ 308] Habitat : Forest and grassland[ 309] Diet : Insects[ 306]
VU
Unknown [ 309]
Corbet's forest shrew
S. corbeti Hutterer , 2009
West-central Africa
Size : 4โ10 cm (2โ4 in) long, plus 4โ9 cm (2โ4 in) tail[ 274] Habitat : Forest[ 310] Diet : Insects[ 306]
DD
Unknown [ 310]
Dudu Akaibe's pygmy shrew
S. akaibei Mukinzi , Hutterer , & Barriere , 2009
Democratic Republic of the Congo
Size : 4โ10 cm (2โ4 in) long, plus 4โ9 cm (2โ4 in) tail[ 274] Habitat : Forest[ 311] Diet : Insects[ 306]
DD
Unknown [ 311]
Grant's forest shrew
S. granti Thomas , 1907
S. g. granti
S. g. mundus
Central Africa
Size : 4โ7 cm (2โ3 in) long, plus 4โ7 cm (2โ3 in) tail[ 312] Habitat : Forest[ 313] Diet : Insects[ 306]
LC
Unknown [ 313]
Greater forest shrew
S. ollula Thomas , 1913
Central Africa
Size : 5โ8 cm (2โ3 in) long, plus 1โ2 cm (0โ1 in) tail[ 314] Habitat : Forest[ 315] Diet : Insects[ 306]
LC
Unknown [ 315]
Howell's forest shrew
S. howelli Jenkins , 1984
Tanzania
Size : 5โ8 cm (2โ3 in) long, plus 3โ5 cm (1โ2 in) tail[ 316] Habitat : Forest[ 317] Diet : Insects[ 306]
LC
Unknown [ 317]
Johnston's forest shrew
S. johnstoni (Dobson , 1888)
West-central and central Africa
Size : 7โ10 cm (3โ4 in) long, plus 4โ7 cm (2โ3 in) tail[ 308] Habitat : Forest[ 318] Diet : Insects[ 306]
LC
Unknown [ 318]
Kongana shrew
S. konganensis Ray & Hutterer , 1996
Central Africa
Size : 4โ10 cm (2โ4 in) long, plus 4โ9 cm (2โ4 in) tail[ 274] Habitat : Forest[ 319] Diet : Insects[ 306]
DD
Unknown [ 319]
Lesser forest shrew
S. oriundus Hollister , 1916
Central Africa
Size : 4โ10 cm (2โ4 in) long, plus 4โ9 cm (2โ4 in) tail[ 274] Habitat : Forest[ 320] Diet : Insects[ 306]
DD
Unknown [ 320]
Moon forest shrew
S. lunaris Thomas , 1906
Central Africa
Size : 7โ10 cm (3โ4 in) long, plus 4โ6 cm (2 in) tail[ 321] Habitat : Forest[ 322] Diet : Insects[ 306]
NT
Unknown [ 322]
Mount Cameroon forest shrew
S. morio (Gray , 1862)
West-central Africa
Size : 6โ8 cm (2โ3 in) long, plus 5โ6 cm (2 in) tail[ 323] Habitat : Forest[ 324] Diet : Insects[ 306]
EN
Unknown [ 324]
Rain forest shrew
S. pluvialis Hutterer & Schlitter , 1996
West-central Africa
Size : 6โ8 cm (2โ3 in) long, plus 5โ8 cm (2โ3 in) tail[ 321] Habitat : Forest[ 325] Diet : Insects[ 306]
DD
Unknown [ 325]
Volcano shrew
S. vulcanorum Hutterer & Verheyan , 1985
Central Africa
Size : 4โ6 cm (2 in) long, plus 4โ7 cm (2โ3 in) tail[ 308] Habitat : Forest and inland wetlands[ 326] Diet : Insects[ 306]
LC
Unknown [ 326]
References
^ a b Nowak , p. 224
^ Jenkins, P. D.; Lunde, D. P.; Moncrieff, C. B. (2009). "Descriptions of New Species of Crocidura (Soricomorpha: Soricidae) from Mainland Southeast Asia, with Synopses of Previously Described Species and Remarks on Biogeography" (PDF) . Bulletin of the American Museum of Natural History . 331 : 356โ405. doi :10.1206/582-10.1 . S2CID 84803490 . Archived (PDF) from the original on 2020-03-13. Retrieved 2024-04-12 .
^ "Fossilworks: Crocidurinae" . Paleobiology Database . University of WisconsinโMadison . Archived from the original on June 6, 2023. Retrieved April 11, 2024 .
^ Nations, J. A.; Giarla, T. C.; Morni, M. A.; William Dee, J.; Swanson, M. T.; Hiller, A. E.; Khan, F. A. A.; Esselstyn, J. A. (2022). "Molecular data from the holotype of the enigmatic Bornean Black Shrew, Suncus ater Medway, 1965 (Soricidae, Crocidurinae), place it in the genus Palawanosorex " . ZooKeys (1137): 17โ31. Bibcode :2022ZooK.1137...17N . doi :10.3897/zookeys.1137.94217 . PMC 9836656 . PMID 36760481 .
^ Wilson, Reeder , pp. 224-262
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^ a b Temple, H. (2017) [errata version of 2016 assessment]. "Crocidura shantungensis " . IUCN Red List of Threatened Species . 2016 : e.T5617A115077696. doi :10.2305/IUCN.UK.2016-3.RLTS.T5617A22300721.en .
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^ a b Dando, T.; Kennerley, R. (2019). "Crocidura yaldeni " . IUCN Red List of Threatened Species . 2019 : e.T112517512A112517515. doi :10.2305/IUCN.UK.2019-1.RLTS.T112517512A112517515.en .
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^ a b Raese, Joshua (2004). "Crocidura leucodon " . Animal Diversity Web . University of Michigan . Archived from the original on April 1, 2023. Retrieved March 30, 2024 .
^ a b Shenbrot, G.; Hutterer, R.; Kryลกtufek, B.; Yigit, N.; Mitsainas, G.; Palomo, L. (2021) [amended version of 2016 assessment]. "Crocidura leucodon " . IUCN Red List of Threatened Species . 2021 : e.T29651A197500630. doi :10.2305/IUCN.UK.2021-1.RLTS.T29651A197500630.en .
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^ a b Hutterer, R. (2016). "Crocidura nigricans " . IUCN Red List of Threatened Species . 2016 : e.T41345A22310112. doi :10.2305/IUCN.UK.2016-2.RLTS.T41345A22310112.en .
^ a b Cassola, F. (2016). "Crocidura foetida " . IUCN Red List of Threatened Species . 2016 : e.T136578A22310170. doi :10.2305/IUCN.UK.2016-2.RLTS.T136578A22310170.en .
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^ Kingdon , p. 63
^ Kingdon , p. 96
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura littoralis " . IUCN Red List of Threatened Species . 2016 : e.T41330A115178517. doi :10.2305/IUCN.UK.2016-3.RLTS.T41330A22305865.en .
^ a b Kennerley, R. (2018). "Crocidura picea " . IUCN Red List of Threatened Species . 2018 : e.T5636A22301535. doi :10.2305/IUCN.UK.2018-1.RLTS.T5636A22301535.en .
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^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura caspica " . IUCN Red List of Threatened Species . 2016 : e.T136444A115207731. doi :10.2305/IUCN.UK.2016-3.RLTS.T136444A22297843.en .
^ a b Johnston, C.; Smith, A. T. (2016). "Crocidura rapax " . IUCN Red List of Threatened Species . 2016 : e.T136783A22294279. doi :10.2305/IUCN.UK.2016-1.RLTS.T136783A22294279.en .
^ a b Woinarski, J.; Burbidge, A. A.; Lumsden, L. (2016). "Crocidura trichura " . IUCN Red List of Threatened Species . 2016 : e.T136379A22304640. doi :10.2305/IUCN.UK.2016-3.RLTS.T136379A22304640.en .
^ Kingdon , p. 65
^ a b Hutterer, R.; Granjon, L. (2017) [errata version of 2016 assessment]. "Crocidura cinderella " . IUCN Red List of Threatened Species . 2016 : e.T41317A115177629. doi :10.2305/IUCN.UK.2016-3.RLTS.T41317A22307375.en .
^ a b Hutterer, R. (2016). "Crocidura congobelgica " . IUCN Red List of Threatened Species . 2016 : e.T5561A22305663. doi :10.2305/IUCN.UK.2016-2.RLTS.T5561A22305663.en .
^ a b Jenkins, P.; Dando, T.; Kennerley, R. (2017). "Crocidura cranbrooki " . IUCN Red List of Threatened Species . 2017 : e.T48296479A48296547. doi :10.2305/IUCN.UK.2017-3.RLTS.T48296479A48296547.en .
^ a b Mitsainas, G.; Vohralรญk, V.; Kennerley, R. (2019). "Crocidura zimmermanni " . IUCN Red List of Threatened Species . 2019 : e.T5588A90818329. doi :10.2305/IUCN.UK.2019-1.RLTS.T5588A90818329.en .
^ Kingdon , p. 68
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura crossei " . IUCN Red List of Threatened Species . 2016 : e.T40624A115175904. doi :10.2305/IUCN.UK.2016-3.RLTS.T40624A22296115.en .
^ Kingdon , p. 56
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^ Kingdon , p. 70
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^ Kingdon , p. 133
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^ Kingdon , p. 71
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^ a b Gerrie, R.; Kennerley, R. (2016). "Crocidura dhofarensis " . IUCN Red List of Threatened Species . 2016 : e.T5599A22303111. doi :10.2305/IUCN.UK.2016-3.RLTS.T5599A22303111.en .
^ a b Gerrie, R.; Kennerley, R. (2016). "Crocidura douceti " . IUCN Red List of Threatened Species . 2016 : e.T40629A22295053. doi :10.2305/IUCN.UK.2016-3.RLTS.T40629A22295053.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura dsinezumi " . IUCN Red List of Threatened Species . 2016 : e.T40627A115176222. doi :10.2305/IUCN.UK.2016-3.RLTS.T40627A22294822.en .
^ Kingdon , p. 57
^ a b Dando, T. (2021). "Crocidura allex " . IUCN Red List of Threatened Species . 2021 : e.T5620A22301002. doi :10.2305/IUCN.UK.2021-2.RLTS.T5620A22301002.en .
^ Kingdon , p. 128
^ a b Kryลกtufek, B.; Kennerley, R. (2019). "Crocidura religiosa " . IUCN Red List of Threatened Species . 2019 : e.T5616A114077410. doi :10.2305/IUCN.UK.2019-1.RLTS.T5616A114077410.en .
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^ a b Cassola, F. (2016). "Crocidura elongata " . IUCN Red List of Threatened Species . 2016 : e.T40630A22294921. doi :10.2305/IUCN.UK.2016-2.RLTS.T40630A22294921.en .
^ a b Cerรญaco, L.; Dando, T.; Kennerley, R. (2019). "Crocidura fingui " . IUCN Red List of Threatened Species . 2019 : e.T111739377A111739380. doi :10.2305/IUCN.UK.2019-1.RLTS.T111739377A111739380.en .
^ Kingdon , p. 76
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^ Kingdon , p. 78
^ Kingdon , p. 79
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^ a b c Kingdon , p. 126
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^ a b Kingdon , p. 84
^ a b Decher, J.; Dando, T.; Kennerley, R.; Hutterer, R. (2019). "Crocidura goliath " . IUCN Red List of Threatened Species . 2019 : e.T112518189A113974008. doi :10.2305/IUCN.UK.2019-3.RLTS.T112518189A113974008.en .
^ Kingdon , p. 107
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^ Kingdon , p. 86
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura grassei " . IUCN Red List of Threatened Species . 2016 : e.T5567A115074501. doi :10.2305/IUCN.UK.2016-3.RLTS.T5567A22305289.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura grandis " . IUCN Red List of Threatened Species . 2016 : e.T5601A115076764. doi :10.2305/IUCN.UK.2016-3.RLTS.T5601A22299006.en .
^ Kingdon , p. 77
^ a b Cassola, F. (2016). "Crocidura flavescens " . IUCN Red List of Threatened Species . 2016 : e.T5628A22300161. doi :10.2305/IUCN.UK.2016-2.RLTS.T5628A22300161.en .
^ a b Knoll, Amanda (2009). "Crocidura russula " . Animal Diversity Web . University of Michigan . Archived from the original on March 14, 2023. Retrieved March 30, 2024 .
^ a b Aulagnier, S.; Hutterer, R.; Kryลกtufek, B.; Yigit, N.; Mitsainas, G.; Palomo, L. (2021) [amended version of 2016 assessment]. "Crocidura russula " . IUCN Red List of Threatened Species . 2021 : e.T29652A197503499. doi :10.2305/IUCN.UK.2021-1.RLTS.T29652A197503499.en .
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^ a b Laginha Pinto Correia, D.; Clayton, E. (2016). "Crocidura wuchihensis " . IUCN Red List of Threatened Species . 2016 : e.T136382A22304269. doi :10.2305/IUCN.UK.2016-1.RLTS.T136382A22304269.en .
^ a b Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura harenna " . IUCN Red List of Threatened Species . 2016 : e.T5633A115078795. doi :10.2305/IUCN.UK.2016-3.RLTS.T5633A22301820.en .
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^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura erica " . IUCN Red List of Threatened Species . 2016 : e.T5626A115078377. doi :10.2305/IUCN.UK.2016-3.RLTS.T5626A22299982.en .
^ Kingdon , p. 89
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^ a b Chiozza, F. (2016). "Crocidura hilliana " . IUCN Red List of Threatened Species . 2016 : e.T136226A22295171. doi :10.2305/IUCN.UK.2016-1.RLTS.T136226A22295171.en .
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^ a b Lunde, D. P.; Ruedas, L. A.; van Strien, N. J. (2020) [errata version of 2017 assessment]. "Crocidura hutanis " . IUCN Red List of Threatened Species . 2017 : e.T136781A165117820. doi :10.2305/IUCN.UK.2017-2.RLTS.T136781A165117820.en .
^ a b Cassola, F. (2016). "Crocidura indochinensis " . IUCN Red List of Threatened Species . 2016 : e.T136733A22299360. doi :10.2305/IUCN.UK.2016-2.RLTS.T136733A22299360.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura susiana " . IUCN Red List of Threatened Species . 2016 : e.T5618A115077828. doi :10.2305/IUCN.UK.2016-3.RLTS.T5618A22300814.en .
^ a b Hutterer, R.; Amori, G.; Kryลกtufek, B. (2019). "Crocidura arispa " . IUCN Red List of Threatened Species . 2019 : e.T136270A22296424. doi :10.2305/IUCN.UK.2019-1.RLTS.T136270A22296424.en .
^ Kingdon , p. 91
^ a b Oguge, N.; Hutterer, R.; Howell, K. (2017) [errata version of 2016 assessment]. "Crocidura jacksoni " . IUCN Red List of Threatened Species . 2016 : e.T41325A115178261. doi :10.2305/IUCN.UK.2016-3.RLTS.T41325A22308229.en .
^ a b Demos, T.; Kennerley, R. (2018). "Crocidura umbra " . IUCN Red List of Threatened Species . 2018 : e.T112465479A112465482. doi :10.2305/IUCN.UK.2018-1.RLTS.T112465479A112465482.en .
^ a b Chiozza, F. (2017) [errata version of 2016 assessment]. "Crocidura maxi " . IUCN Red List of Threatened Species . 2016 : e.T41336A115179357. doi :10.2305/IUCN.UK.2016-3.RLTS.T41336A22306419.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura jenkinsi " . IUCN Red List of Threatened Species . 2016 : e.T5604A115077051. doi :10.2305/IUCN.UK.2016-3.RLTS.T5604A22299101.en .
^ Kingdon , p. 92
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^ a b Molur, S. (2016). "Crocidura pullata " . IUCN Red List of Threatened Species . 2016 : e.T41353A22311057. doi :10.2305/IUCN.UK.2016-1.RLTS.T41353A22311057.en .
^ a b Gerrie, R.; Kennerley, R. (2017). "Crocidura katinka " . IUCN Red List of Threatened Species . 2017 : e.T136634A22303188. doi :10.2305/IUCN.UK.2017-2.RLTS.T136634A22303188.en .
^ a b Dando, T.; Kennerley, R. (2018) [errata version of 2017 assessment]. "Crocidura monax " . IUCN Red List of Threatened Species . 2017 : e.T111767309A123795933. doi :10.2305/IUCN.UK.2017-3.RLTS.T111767309A22304558.en .
^ a b Clayton, E.; Kennerley, R. (2016). "Crocidura baluensis " . IUCN Red List of Threatened Species . 2016 : e.T136726A22300422. doi :10.2305/IUCN.UK.2016-3.RLTS.T136726A22300422.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura lanosa " . IUCN Red List of Threatened Species . 2016 : e.T5569A115074754. doi :10.2305/IUCN.UK.2016-3.RLTS.T5569A22303856.en .
^ a b Kingdon , p. 93
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura kivuana " . IUCN Red List of Threatened Species . 2016 : e.T5568A115074630. doi :10.2305/IUCN.UK.2016-3.RLTS.T5568A22305202.en .
^ a b Hutterer, R. (2016). "Crocidura lamottei " . IUCN Red List of Threatened Species . 2016 : e.T41326A22308096. doi :10.2305/IUCN.UK.2016-2.RLTS.T41326A22308096.en .
^ Kingdon , p. 85
^ a b Hutterer, R. (2008). "Crocidura grandiceps " . IUCN Red List of Threatened Species . 2008 : e.T41321A10443138. doi :10.2305/IUCN.UK.2008.RLTS.T41321A10443138.en .
^ a b Hutterer, R. (2016). "Crocidura latona " . IUCN Red List of Threatened Species . 2016 : e.T5570A22303960. doi :10.2305/IUCN.UK.2016-2.RLTS.T5570A22303960.en .
^ a b Cassola, F. (2016). "Crocidura watasei " . IUCN Red List of Threatened Species . 2016 : e.T136341A22310365. doi :10.2305/IUCN.UK.2016-2.RLTS.T136341A22310365.en .
^ Kingdon , p. 132
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura silacea " . IUCN Red List of Threatened Species . 2016 : e.T41357A115181391. doi :10.2305/IUCN.UK.2016-3.RLTS.T41357A22311376.en .
^ Kingdon , p. 90
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura hirta " . IUCN Red List of Threatened Species . 2016 : e.T41323A115178068. doi :10.2305/IUCN.UK.2016-3.RLTS.T41323A22307953.en .
^ a b Hutterer, R. (2016). "Crocidura serezkyensis " . IUCN Red List of Threatened Species . 2016 : e.T29654A22296786. doi :10.2305/IUCN.UK.2016-2.RLTS.T29654A22296786.en .
^ a b Kryลกtufek, B.; Gazzard, A. (2023). "Crocidura suaveolens " . IUCN Red List of Threatened Species . 2023 : e.T221735459A221735673. doi :10.2305/IUCN.UK.2023-1.RLTS.T221735459A221735673.en .
^ Kingdon , p. 72
^ a b Hutterer, R.; Howell, K.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura dolichura " . IUCN Red List of Threatened Species . 2016 : e.T40628A115176367. doi :10.2305/IUCN.UK.2016-3.RLTS.T40628A22294674.en .
^ Kingdon , p. 98
^ a b Kennerley, R.; Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura lucina " . IUCN Red List of Threatened Species . 2016 : e.T5572A115075017. doi :10.2305/IUCN.UK.2016-3.RLTS.T5572A22304095.en .
^ Kingdon , p. 99
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura ludia " . IUCN Red List of Threatened Species . 2016 : e.T5589A115075820. doi :10.2305/IUCN.UK.2016-3.RLTS.T5589A22302289.en .
^ a b Heaney, L.; Tabaranza, B. (2016). "Crocidura grayi " . IUCN Red List of Threatened Species . 2016 : e.T5602A22298900. doi :10.2305/IUCN.UK.2016-2.RLTS.T5602A22298900.en .
^ a b Plumptre, A. J.; Dando, T.; Kennerley, R. (2020) [amended version of 2019 assessment]. "Crocidura lwiroensis " . IUCN Red List of Threatened Species . 2020 : e.T112139630A166620647. doi :10.2305/IUCN.UK.2020-1.RLTS.T112139630A166620647.en .
^ a b Kennerley, R.; Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura macmillani " . IUCN Red List of Threatened Species . 2016 : e.T5634A115078974. doi :10.2305/IUCN.UK.2016-3.RLTS.T5634A22301723.en .
^ a b Kingdon , p. 104
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura maquassiensis " . IUCN Red List of Threatened Species . 2016 : e.T5576A115075148. doi :10.2305/IUCN.UK.2016-3.RLTS.T5576A22303779.en .
^ a b Cassola, F. (2016). "Crocidura malayana " . IUCN Red List of Threatened Species . 2016 : e.T5605A22298569. doi :10.2305/IUCN.UK.2016-2.RLTS.T5605A22298569.en .
^ a b Kennerley, R. (2016). "Crocidura virgata " . IUCN Red List of Threatened Species . 2016 : e.T136602A22309454. doi :10.2305/IUCN.UK.2016-3.RLTS.T136602A22309454.en .
^ a b Demos, T.; Kennerley, R. (2019). "Crocidura manengubae " . IUCN Red List of Threatened Species . 2019 : e.T5575A22303668. doi :10.2305/IUCN.UK.2019-1.RLTS.T5575A22303668.en .
^ a b Kingdon , p. 100
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura lusitania " . IUCN Red List of Threatened Species . 2016 : e.T41332A115178805. doi :10.2305/IUCN.UK.2016-3.RLTS.T41332A22306053.en .
^ a b Giarla, T.; Dando, T.; Kennerley, R. (2017). "Crocidura mdumai " . IUCN Red List of Threatened Species . 2017 : e.T112503346A112503349. doi :10.2305/IUCN.UK.2017-3.RLTS.T112503346A112503349.en .
^ a b Heaney, L. (2016). "Crocidura beatus " . IUCN Red List of Threatened Species . 2016 : e.T5597A22302943. doi :10.2305/IUCN.UK.2016-2.RLTS.T5597A22302943.en .
^ a b Gerrie, R.; Kennerley, R. (2016). "Crocidura mindorus " . IUCN Red List of Threatened Species . 2016 : e.T48269124A54823641. doi :10.2305/IUCN.UK.2016-3.RLTS.T48269124A54823641.en .
^ a b Hutterer, R. (2019). "Crocidura montis " . IUCN Red List of Threatened Species . 2019 : e.T41338A22306657. doi :10.2305/IUCN.UK.2019-1.RLTS.T41338A22306657.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura luna " . IUCN Red List of Threatened Species . 2016 : e.T41331A115178646. doi :10.2305/IUCN.UK.2016-3.RLTS.T41331A22305942.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura musseri " . IUCN Red List of Threatened Species . 2016 : e.T136711A115211595. doi :10.2305/IUCN.UK.2016-3.RLTS.T136711A22298834.en .
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura nimbasilvanus " . IUCN Red List of Threatened Species . 2019 : e.T112519468A112519471. doi :10.2305/IUCN.UK.2019-3.RLTS.T112519468A112519471.en .
^ a b Dando, T.; Kennerley, R. (2020) [amended version of 2019 assessment]. "Crocidura munissii " . IUCN Red List of Threatened Species . 2020 : e.T112503202A166619293. doi :10.2305/IUCN.UK.2020-1.RLTS.T112503202A166619293.en .
^ a b Kingdon , p. 134
^ a b Kerbis Peterhans, J.; Dieterlen, F.; Hutterer, R. (2008). "Crocidura stenocephala " . IUCN Red List of Threatened Species . 2008 : e.T5582A11371843. doi :10.2305/IUCN.UK.2008.RLTS.T5582A11371843.en .
^ a b Hutterer, R.; Shenbrot, G. (2017). "Crocidura ramona " . IUCN Red List of Threatened Species . 2017 : e.T136722A89475013. doi :10.2305/IUCN.UK.2017-2.RLTS.T136722A89475013.en .
^ a b Clayton, E.; Kennerley, R. (2018). "Crocidura negrina " . IUCN Red List of Threatened Species . 2018 : e.T5609A22298060. doi :10.2305/IUCN.UK.2018-1.RLTS.T5609A22298060.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura nicobarica " . IUCN Red List of Threatened Species . 2016 : e.T5610A115077236. doi :10.2305/IUCN.UK.2016-3.RLTS.T5610A22297919.en .
^ Kingdon , p. 113
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura nigeriae " . IUCN Red List of Threatened Species . 2016 : e.T41344A115179933. doi :10.2305/IUCN.UK.2016-3.RLTS.T41344A22307167.en .
^ a b Hutterer, R. (2008). "Crocidura nimbae " . IUCN Red List of Threatened Species . 2008 : e.T5578A11354307. doi :10.2305/IUCN.UK.2008.RLTS.T5578A11354307.en .
^ Kingdon , p. 117
^ a b Gerrie, R.; Kennerley, R. (2017). "Crocidura niobe " . IUCN Red List of Threatened Species . 2017 : e.T41347A22309898. doi :10.2305/IUCN.UK.2017-2.RLTS.T41347A22309898.en .
^ a b Amori, G.; Hutterer, R.; Kryลกtufek, B.; Yigit, N. (2022). "Crocidura pachyura " . IUCN Red List of Threatened Species . 2022 : e.T40623A22296091. doi :10.2305/IUCN.UK.2022-2.RLTS.T40623A22296091.en .
^ Kingdon , p. 103
^ a b Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura macowi " . IUCN Red List of Threatened Species . 2016 : e.T5635A115079098. doi :10.2305/IUCN.UK.2016-3.RLTS.T5635A22301650.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura orientalis " . IUCN Red List of Threatened Species . 2016 : e.T5611A115077427. doi :10.2305/IUCN.UK.2016-3.RLTS.T5611A22298371.en .
^ a b Cassola, F. (2016). "Crocidura palawanensis " . IUCN Red List of Threatened Species . 2016 : e.T5613A22297590. doi :10.2305/IUCN.UK.2016-2.RLTS.T5613A22297590.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura pergrisea " . IUCN Red List of Threatened Species . 2016 : e.T5615A115077570. doi :10.2305/IUCN.UK.2016-3.RLTS.T5615A22297761.en .
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura panayensis " . IUCN Red List of Threatened Species . 2019 : e.T48297786A48297816. doi :10.2305/IUCN.UK.2019-1.RLTS.T48297786A48297816.en .
^ a b Ruedas, L. (2016). "Crocidura negligens " . IUCN Red List of Threatened Species . 2016 : e.T136786A22296214. doi :10.2305/IUCN.UK.2016-1.RLTS.T136786A22296214.en .
^ a b Engelbrektsson, P.; Kennerley, R. (2016). "Crocidura phanluongi " . IUCN Red List of Threatened Species . 2016 : e.T45954204A45973011. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954204A45973011.en .
^ a b Engelbrektsson, P. (2016). "Crocidura phuquocensis " . IUCN Red List of Threatened Species . 2016 : e.T45954289A45973016. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954289A45973016.en .
^ Kingdon , p. 124
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura pitmani " . IUCN Red List of Threatened Species . 2016 : e.T5637A115079206. doi :10.2305/IUCN.UK.2016-3.RLTS.T5637A22301462.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura polia " . IUCN Red List of Threatened Species . 2016 : e.T5591A115075965. doi :10.2305/IUCN.UK.2016-3.RLTS.T5591A22302474.en .
^ Kingdon , p. 127
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura raineyi " . IUCN Red List of Threatened Species . 2016 : e.T5592A115076066. doi :10.2305/IUCN.UK.2016-3.RLTS.T5592A22302388.en .
^ Kingdon , p. 69
^ a b c Baxter, R.; Hutterer, R.; Griffin, M.; Howell, K. (2017) [errata version of 2016 assessment]. "Crocidura cyanea " . IUCN Red List of Threatened Species . 2016 : e.T40625A115176043. doi :10.2305/IUCN.UK.2016-3.RLTS.T40625A22294530.en .
^ a b Hutterer, R.; Kerbis Peterhans, J. (2017) [errata version of 2016 assessment]. "Crocidura roosevelti " . IUCN Red List of Threatened Species . 2016 : e.T41355A115181119. doi :10.2305/IUCN.UK.2016-3.RLTS.T41355A22310848.en .
^ a b Laginha Pinto Correia, D. (2016). "Crocidura orii " . IUCN Red List of Threatened Species . 2016 : e.T5590A22302169. doi :10.2305/IUCN.UK.2016-1.RLTS.T5590A22302169.en .
^ a b Jenkins, P.; Dando, T.; Kennerley, R. (2019). "Crocidura sapaensis " . IUCN Red List of Threatened Species . 2019 : e.T112465460A112465464. doi :10.2305/IUCN.UK.2019-1.RLTS.T112465460A112465464.en .
^ a b Aulagnier, S.; Gerrie, R.; Kennerley, R. (2019). "Crocidura tarfayensis " . IUCN Red List of Threatened Species . 2019 : e.T41361A22308881. doi :10.2305/IUCN.UK.2019-1.RLTS.T41361A22308881.en .
^ Kingdon , p. 121
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura pasha " . IUCN Red List of Threatened Species . 2016 : e.T41350A115180721. doi :10.2305/IUCN.UK.2016-3.RLTS.T41350A22310542.en .
^ Kingdon , p. 112
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura nanilla " . IUCN Red List of Threatened Species . 2016 : e.T41342A115179768. doi :10.2305/IUCN.UK.2016-3.RLTS.T41342A97250815.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura viaria " . IUCN Red List of Threatened Species . 2016 : e.T41364A115181785. doi :10.2305/IUCN.UK.2016-3.RLTS.T41364A22308708.en .
^ a b Hutterer, R.; Jenkins, P. (2017) [errata version of 2016 assessment]. "Crocidura fulvastra " . IUCN Red List of Threatened Species . 2016 : e.T40632A115176689. doi :10.2305/IUCN.UK.2016-3.RLTS.T40632A22295294.en .
^ Kingdon , p. 97
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura longipes " . IUCN Red List of Threatened Species . 2016 : e.T5571A115074897. doi :10.2305/IUCN.UK.2016-3.RLTS.T5571A22304019.en .
^ a b Stubbe, M.; Samiya, R.; Ariunbold, J.; Buuveibaatar, V.; Dorjderem, S.; Monkhzul, Ts.; Otgonbaatar, M.; Tsogbadrakh, M. (2017) [errata version of 2016 assessment]. "Crocidura sibirica " . IUCN Red List of Threatened Species . 2016 : e.T41356A115181252. doi :10.2305/IUCN.UK.2016-3.RLTS.T41356A22310743.en .
^ a b Dando, T.; Kennerley, R. (2020) [amended version of 2019 assessment]. "Crocidura ninoyi " . IUCN Red List of Threatened Species . 2020 : e.T48269338A166612644. doi :10.2305/IUCN.UK.2020-1.RLTS.T48269338A166612644.en .
^ a b Giovanni Amori. (2016). "Crocidura sicula " . IUCN Red List of Threatened Species . 2016 : e.T29655A2791535. doi :10.2305/IUCN.UK.2016-2.RLTS.T29655A2791535.en .
^ a b Meegaskumbura, S.; Meegaskumbura, M. (2008). "Crocidura hikmiya " . IUCN Red List of Threatened Species . 2008 : e.T136596A4316355. doi :10.2305/IUCN.UK.2008.RLTS.T136596A4316355.en .
^ Kingdon , p. 120
^ a b Kennerley, R. (2016). "Crocidura fumosa " . IUCN Red List of Threatened Species . 2016 : e.T5630A22299610. doi :10.2305/IUCN.UK.2016-3.RLTS.T5630A22299610.en .
^ a b Engelbrektsson, P. (2016). "Crocidura sokolovi " . IUCN Red List of Threatened Species . 2016 : e.T45954331A45973021. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954331A45973021.en .
^ a b Kingdon , p. 111
^ a b Cassola, F. (2019). "Crocidura nana " . IUCN Red List of Threatened Species . 2019 : e.T41341A22306927. doi :10.2305/IUCN.UK.2019-1.RLTS.T41341A22306927.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura somalica " . IUCN Red List of Threatened Species . 2016 : e.T41359A115181668. doi :10.2305/IUCN.UK.2016-3.RLTS.T41359A22311245.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura fuliginosa " . IUCN Red List of Threatened Species . 2016 : e.T40631A115176525. doi :10.2305/IUCN.UK.2016-3.RLTS.T40631A22295396.en .
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura miya " . IUCN Red List of Threatened Species . 2019 : e.T5608A22298694. doi :10.2305/IUCN.UK.2019-3.RLTS.T5608A22298694.en .
^ a b Chiozza, F. (2016). "Crocidura lea " . IUCN Red List of Threatened Species . 2016 : e.T41328A22308315. doi :10.2305/IUCN.UK.2016-2.RLTS.T41328A22308315.en .
^ a b Chiozza, F. (2016). "Crocidura levicula " . IUCN Red List of Threatened Species . 2016 : e.T41329A22305755. doi :10.2305/IUCN.UK.2016-2.RLTS.T41329A22305755.en .
^ a b Cassola, F. (2016). "Crocidura rhoditis " . IUCN Red List of Threatened Species . 2016 : e.T41354A22310945. doi :10.2305/IUCN.UK.2016-2.RLTS.T41354A22310945.en .
^ a b Cassola, F. (2016). "Crocidura lepidura " . IUCN Red List of Threatened Species . 2016 : e.T136792A22296335. doi :10.2305/IUCN.UK.2016-2.RLTS.T136792A22296335.en .
^ a b Cassola, F. (2016). "Crocidura paradoxura " . IUCN Red List of Threatened Species . 2016 : e.T5614A22297493. doi :10.2305/IUCN.UK.2016-2.RLTS.T5614A22297493.en .
^ a b Chiozza, F. (2016). "Crocidura monticola " . IUCN Red List of Threatened Species . 2016 : e.T41337A22306541. doi :10.2305/IUCN.UK.2016-2.RLTS.T41337A22306541.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura mariquensis " . IUCN Red List of Threatened Species . 2016 : e.T41334A115179070. doi :10.2305/IUCN.UK.2016-3.RLTS.T41334A22306233.en .
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura thomensis " . IUCN Red List of Threatened Species . 2016 : e.T5594A115076316. doi :10.2305/IUCN.UK.2016-3.RLTS.T5594A22303259.en .
^ a b Clayton, E. (2016). "Crocidura tanakae " . IUCN Red List of Threatened Species . 2016 : e.T136238A22294077. doi :10.2305/IUCN.UK.2016-2.RLTS.T136238A22294077.en .
^ Kingdon , p. 135
^ a b Dando, T.; Kennerley, R. (2019). "Crocidura tansaniana " . IUCN Red List of Threatened Species . 2019 : e.T112513345A112705120. doi :10.2305/IUCN.UK.2019-2.RLTS.T112513345A112705120.en .
^ a b Kennerley, R. (2016). "Crocidura tarella " . IUCN Red List of Threatened Species . 2016 : e.T41360A22311139. doi :10.2305/IUCN.UK.2016-3.RLTS.T41360A22311139.en .
^ a b Kingdon , p. 138
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura telfordi " . IUCN Red List of Threatened Species . 2016 : e.T5584A115075552. doi :10.2305/IUCN.UK.2016-3.RLTS.T5584A22304467.en .
^ a b Lavrenchenko, L. (2017) [errata version of 2016 assessment]. "Crocidura thalia " . IUCN Red List of Threatened Species . 2016 : e.T5585A115075702. doi :10.2305/IUCN.UK.2016-3.RLTS.T5585A22302660.en .
^ Kingdon , p. 139
^ a b Hutterer, R. (2016). "Crocidura theresae " . IUCN Red List of Threatened Species . 2016 : e.T41362A22308991. doi :10.2305/IUCN.UK.2016-2.RLTS.T41362A22308991.en .
^ a b Demos, T.; Kennerley, R. (2019). "Crocidura brunnea " . IUCN Red List of Threatened Species . 2019 : e.T136742A22297393. doi :10.2305/IUCN.UK.2019-1.RLTS.T136742A22297393.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura tenuis " . IUCN Red List of Threatened Species . 2016 : e.T5619A115077931. doi :10.2305/IUCN.UK.2016-3.RLTS.T5619A22300896.en .
^ Kingdon , p. 140
^ a b Hutterer, R. (2016). "Crocidura turba " . IUCN Red List of Threatened Species . 2016 : e.T41363A22308481. doi :10.2305/IUCN.UK.2016-2.RLTS.T41363A22308481.en .
^ Kingdon , p. 131
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura selina " . IUCN Red List of Threatened Species . 2016 : e.T5593A115076180. doi :10.2305/IUCN.UK.2016-3.RLTS.T5593A22303358.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura mutesae " . IUCN Red List of Threatened Species . 2016 : e.T41340A115179654. doi :10.2305/IUCN.UK.2016-3.RLTS.T41340A22306859.en .
^ Kingdon , p. 141
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura ultima " . IUCN Red List of Threatened Species . 2016 : e.T5638A115079307. doi :10.2305/IUCN.UK.2016-3.RLTS.T5638A22301388.en .
^ Kingdon , p. 149
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Crocidura zimmeri " . IUCN Red List of Threatened Species . 2016 : e.T5640A115079422. doi :10.2305/IUCN.UK.2016-3.RLTS.T5640A22301128.en .
^ Kingdon , p. 142
^ a b Giarla, T.; Dando, T.; Kennerley, R. (2018) [errata version of 2017 assessment]. "Crocidura usambarae " . IUCN Red List of Threatened Species . 2017 : e.T112515709A123796512. doi :10.2305/IUCN.UK.2017-3.RLTS.T112515709A112705149.en .
^ a b c Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura lasiura " . IUCN Red List of Threatened Species . 2016 : e.T41327A115178387. doi :10.2305/IUCN.UK.2016-3.RLTS.T41327A22308395.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura voi " . IUCN Red List of Threatened Species . 2016 : e.T41365A115182001. doi :10.2305/IUCN.UK.2016-3.RLTS.T41365A22309320.en .
^ a b Chiozza, F. (2016). "Crocidura vorax " . IUCN Red List of Threatened Species . 2016 : e.T136454A22301948. doi :10.2305/IUCN.UK.2016-2.RLTS.T136454A22301948.en .
^ Kingdon , p. 110
^ a b Hutterer, R.; Howell, K. (2017) [errata version of 2016 assessment]. "Crocidura muricauda " . IUCN Red List of Threatened Species . 2016 : e.T41339A115179514. doi :10.2305/IUCN.UK.2016-3.RLTS.T41339A22306757.en .
^ Kingdon , p. 118
^ a b Nicolas, V.; Dando, T.; Kennerley, R. (2018) [errata version of 2017 assessment]. "Crocidura obscurior " . IUCN Red List of Threatened Species . 2017 : e.T111754705A123795806. doi :10.2305/IUCN.UK.2017-3.RLTS.T111754705A111754743.en .
^ Kingdon , p. 145
^ Kingdon , p. 146
^ a b Kingdon , p. 147
^ a b Hutterer, R. (2016). "Crocidura xantippe " . IUCN Red List of Threatened Species . 2016 : e.T5639A22301319. doi :10.2305/IUCN.UK.2016-2.RLTS.T5639A22301319.en .
^ a b Cassola, F. (2017) [errata version of 2016 assessment]. "Crocidura yankariensis " . IUCN Red List of Threatened Species . 2016 : e.T41367A115182133. doi :10.2305/IUCN.UK.2016-3.RLTS.T41367A22309073.en .
^ Kingdon , p. 148
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Crocidura zarudnyi " . IUCN Red List of Threatened Species . 2016 : e.T41369A115182396. doi :10.2305/IUCN.UK.2016-3.RLTS.T41369A22309535.en .
^ a b c Nowak , p. 227
^ a b c Tsytsulina, K.; Formozov, N.; Sheftel, B. (2017) [errata version of 2016 assessment]. "Diplomesodon pulchellum " . IUCN Red List of Threatened Species . 2016 : e.T41448A115186837. doi :10.2305/IUCN.UK.2016-3.RLTS.T41448A22293795.en .
^ a b c Nowak , p. 226
^ a b de A. Goonatilake, W. L. D. P. T. S.; Molur, S. (2020). "Feroculus feroculus " . IUCN Red List of Threatened Species . 2020 : e.T8553A22292753. doi :10.2305/IUCN.UK.2020-2.RLTS.T8553A22292753.en .
^ a b c d e f Nowak , p. 222
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^ a b Gerrie, R.; Kennerley, R. (2019). "Paracrocidura maxima " . IUCN Red List of Threatened Species . 2019 : e.T16043A22311579. doi :10.2305/IUCN.UK.2019-1.RLTS.T16043A22311579.en .
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^ a b c d e f g Nowak , p. 225
^ a b Kerbis Peterhans, J. (2017) [errata version of 2016 assessment]. "Ruwenzorisorex suncoides " . IUCN Red List of Threatened Species . 2016 : e.T19790A115153189. doi :10.2305/IUCN.UK.2016-3.RLTS.T19790A22290635.en .
^ a b Hatchett, Jani (2000). "Scutisorex somereni " . Animal Diversity Web . University of Michigan . Archived from the original on October 31, 2023. Retrieved March 30, 2024 .
^ a b Cassola, F. (2020) [amended version of 2016 assessment]. "Scutisorex somereni " . IUCN Red List of Threatened Species . 2020 : e.T41449A166519064. doi :10.2305/IUCN.UK.2020-1.RLTS.T41449A166519064.en .
^ a b Nowak , pp. 228โ229
^ a b Dando, T.; Kennerley, R. (2019). "Scutisorex thori " . IUCN Red List of Threatened Species . 2019 : e.T112390882A112390898. doi :10.2305/IUCN.UK.2019-1.RLTS.T112390882A112390898.en .
^ a b de A. Goonatilake, W. L. D. P. T. S.; Nameer, P. O.; Molur, S. (2016) [errata version of 2008 assessment]. "Solisorex pearsoni " . IUCN Red List of Threatened Species . 2008 : e.T20332A88692605. doi :10.2305/IUCN.UK.2008.RLTS.T20332A9186854.en .
^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab Kingdon , p. 178
^ a b Molur, S. (2016). "Suncus stoliczkanus " . IUCN Red List of Threatened Species . 2016 : e.T41441A22288889. doi :10.2305/IUCN.UK.2016-2.RLTS.T41441A22288889.en .
^ a b Molur, S.; Nameer, P. O.; de A. Goonatilake, W. I. L. D. P. T. S. (2008). "Suncus montanus " . IUCN Red List of Threatened Species . 2008 : e.T21147A9251556. doi :10.2305/IUCN.UK.2008.RLTS.T21147A9251556.en .
^ a b c Kingdon , p. 175
^ a b Hutterer, R.; Molur, S.; Heaney, L. (2016). "Suncus murinus " . IUCN Red List of Threatened Species . 2016 : e.T41440A22287830. doi :10.2305/IUCN.UK.2016-2.RLTS.T41440A22287830.en .
^ Lench, Jessica (2004). "Suncus murinus " . Animal Diversity Web . University of Michigan . Archived from the original on December 9, 2023. Retrieved March 30, 2024 .
^ a b Chiozza, F. (2016). "Suncus ater " . IUCN Red List of Threatened Species . 2016 : e.T21141A22290051. doi :10.2305/IUCN.UK.2016-2.RLTS.T21141A22290051.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Suncus hosei " . IUCN Red List of Threatened Species . 2016 : e.T21144A115160553. doi :10.2305/IUCN.UK.2016-3.RLTS.T21144A22289712.en .
^ a b Cassola, F. (2017). "Suncus megalura " . IUCN Red List of Threatened Species . 2017 : e.T41446A22288732. doi :10.2305/IUCN.UK.2017-2.RLTS.T41446A22288732.en .
^ a b Molur, S. (2017) [errata version of 2016 assessment]. "Suncus dayi " . IUCN Red List of Threatened Species . 2016 : e.T21142A115160385. doi :10.2305/IUCN.UK.2016-3.RLTS.T21142A22289933.en .
^ a b Aulagnier, S.; Hutterer, R.; Jenkins, P.; Bukhnikashvili, A.; Kryลกtufek, B.; Kock, D. (2017). "Suncus etruscus " . IUCN Red List of Threatened Species . 2017 : e.T90389138A22288134. doi :10.2305/IUCN.UK.2017-2.RLTS.T90389138A22288134.en .
^ Ferry, Anna (2005). "Suncus etruscus " . Animal Diversity Web . University of Michigan . Archived from the original on April 6, 2024. Retrieved March 30, 2024 .
^ a b Clayton, E. (2017) [errata version of 2016 assessment]. "Suncus mertensi " . IUCN Red List of Threatened Species . 2016 : e.T21146A115160697. doi :10.2305/IUCN.UK.2016-3.RLTS.T21146A22289342.en .
^ a b Cassola, F. (2017). "Suncus lixus " . IUCN Red List of Threatened Species . 2017 : e.T41438A22288609. doi :10.2305/IUCN.UK.2017-2.RLTS.T41438A22288609.en .
^ a b Engelbrektsson, P. (2016). "Suncus hututsi " . IUCN Red List of Threatened Species . 2016 : e.T45954392A45973056. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954392A45973056.en .
^ a b Dando, T.; Kennerley, R. (2019). "Suncus zeylanicus " . IUCN Red List of Threatened Species . 2019 : e.T21148A22289100. doi :10.2305/IUCN.UK.2019-3.RLTS.T21148A22289100.en .
^ a b Cassola, F. (2017). "Suncus infinitesimus " . IUCN Red List of Threatened Species . 2017 : e.T41437A22288518. doi :10.2305/IUCN.UK.2017-2.RLTS.T41437A22288518.en .
^ a b Cassola, F. (2017). "Suncus varilla " . IUCN Red List of Threatened Species . 2017 : e.T41442A22288989. doi :10.2305/IUCN.UK.2017-2.RLTS.T41442A22288989.en .
^ a b Laginha Pinto Correia, D. (2016). "Suncus malayanus " . IUCN Red List of Threatened Species . 2016 : e.T21145A22289421. doi :10.2305/IUCN.UK.2016-1.RLTS.T21145A22289421.en .
^ a b Cassola, F. (2017). "Suncus remyi " . IUCN Red List of Threatened Species . 2017 : e.T21140A22289636. doi :10.2305/IUCN.UK.2017-2.RLTS.T21140A22289636.en .
^ a b de A. Goonatilake, W. L. D. P. T. S. (2020). "Suncus fellowesgordoni " . IUCN Red List of Threatened Species . 2020 : e.T21143A22289813. doi :10.2305/IUCN.UK.2020-2.RLTS.T21143A22289813.en .
^ Kingdon , p. 173
^ a b Kennerley, R. (2016). "Suncus aequatorius " . IUCN Red List of Threatened Species . 2016 : e.T136224A22289522. doi :10.2305/IUCN.UK.2016-3.RLTS.T136224A22289522.en .
^ Kingdon , p. 172
^ a b Engelbrektsson, P. (2016). "Sylvisorex silvanorum " . IUCN Red List of Threatened Species . 2016 : e.T45954439A45973071. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954439A45973071.en .
^ a b c d e f g h i j k l m n o Kingdon , p. 186
^ a b Kennerley, R. (2019). "Sylvisorex isabellae " . IUCN Red List of Threatened Species . 2019 : e.T21220A22290742. doi :10.2305/IUCN.UK.2019-3.RLTS.T21220A22290742.en .
^ a b c Kingdon , p. 187
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Sylvisorex camerunensis " . IUCN Red List of Threatened Species . 2016 : e.T45051A115202124. doi :10.2305/IUCN.UK.2016-3.RLTS.T45051A22290959.en .
^ a b Dando, T.; Kennerley, R.; Hutterer, R. (2017). "Sylvisorex corbeti " . IUCN Red List of Threatened Species . 2017 : e.T48294480A48294485. doi :10.2305/IUCN.UK.2017-3.RLTS.T48294480A48294485.en .
^ a b Engelbrektsson, P. (2016). "Sylvisorex akaibei " . IUCN Red List of Threatened Species . 2016 : e.T45954406A45973066. doi :10.2305/IUCN.UK.2016-1.RLTS.T45954406A45973066.en .
^ Kingdon , p. 188
^ a b Hutterer, R. (2016). "Sylvisorex granti " . IUCN Red List of Threatened Species . 2016 : e.T41443A22291731. doi :10.2305/IUCN.UK.2016-2.RLTS.T41443A22291731.en .
^ Kingdon , p. 195
^ a b Cassola, F. (2017). "Sylvisorex ollula " . IUCN Red List of Threatened Species . 2017 : e.T48294467A22291264. doi :10.2305/IUCN.UK.2017-3.RLTS.T48294467A22291264.en .
^ Kingdon , p. 189
^ a b Kennerley, R. (2016). "Sylvisorex howelli " . IUCN Red List of Threatened Species . 2016 : e.T21216A22290854. doi :10.2305/IUCN.UK.2016-3.RLTS.T21216A22290854.en .
^ a b Cassola, F. (2017). "Sylvisorex johnstoni " . IUCN Red List of Threatened Species . 2017 : e.T41444A22291821. doi :10.2305/IUCN.UK.2017-2.RLTS.T41444A22291821.en .
^ a b Gerrie, R.; Kennerley, R. (2016). "Sylvisorex konganensis " . IUCN Red List of Threatened Species . 2016 : e.T45050A22291359. doi :10.2305/IUCN.UK.2016-3.RLTS.T45050A22291359.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Sylvisorex oriundus " . IUCN Red List of Threatened Species . 2016 : e.T21222A115161223. doi :10.2305/IUCN.UK.2016-3.RLTS.T21222A22291447.en .
^ a b Kingdon , p. 193
^ a b Kennerley, R. (2019). "Sylvisorex lunaris " . IUCN Red List of Threatened Species . 2019 : e.T41445A22291627. doi :10.2305/IUCN.UK.2019-1.RLTS.T41445A22291627.en .
^ Kingdon , p. 194
^ a b Kennerley, R. (2017) [errata version of 2016 assessment]. "Sylvisorex morio " . IUCN Red List of Threatened Species . 2016 : e.T21221A115161075. doi :10.2305/IUCN.UK.2016-3.RLTS.T21221A22291520.en .
^ a b Gerrie, R.; Kennerley, R. (2017) [errata version of 2016 assessment]. "Sylvisorex pluvialis " . IUCN Red List of Threatened Species . 2016 : e.T45049A115202001. doi :10.2305/IUCN.UK.2016-3.RLTS.T45049A22291176.en .
^ a b Gerrie, R.; Kennerley, R. (2019). "Sylvisorex vulcanorum " . IUCN Red List of Threatened Species . 2019 : e.T21219A22291074. doi :10.2305/IUCN.UK.2019-1.RLTS.T21219A22291074.en .
Sources
By class By subclass or infraclass By order
Afrosoricids (golden moles, otter shrews, and tenrecs)
Artiodactyls (even-toed ungulates)
Carnivorans (carnivores)
Chiropterans (bats)
Cingulates (armadillos)
Dasyuromorphs (quolls, dunnarts, and other Australian carnivores)
Didelphimorphs (opossums)
Diprotodonts (kangaroos, possums, and wombats)
Eulipotyphlans (hedgehogs, moles, and shrews)
Lagomorphs (hares, rabbits, and pikas)
Macroscelids (elephant shrews)
Peramelemorphs (bandicoots and bilbies)
Perissodactyls (odd-toed ungulates)
Pilosans (anteaters and sloths)
Primates
Rodents
Scandentians (treeshrews)
By suborder or family
By species