Species in the mammal family Viverridae
Four viverrid species (clockwise from top left): the Asian palm civet (Paradoxurus hermaphroditus ), the common genet (Genetta genetta ), the binturong (Arctictis binturong ), and the masked palm civet (Paguma larvata )
Viverridae is a family of mammals in the order Carnivora , composed mainly of the civets and genets . A member of this family is called a viverrid. They are widespread primarily throughout Africa, India, and southeast Asia, and are found primarily in forests, shrublands, and grasslands, though some species can be found in savannas or wetlands. Most viverrids are 40 to 65 cm (16 to 26 in) long, plus a 35 to 60 cm (14 to 24 in) tail, though the West African oyan can be as small as 30 cm (12 in) plus a 35 cm (14 in) tail, and some binturongs can be up to 96 cm (38 in) plus a 89 cm (35 in) tail. Most species do not have population estimates, though three viverrids are classified as endangered , and one, the Malabar large-spotted civet , is classified as critically endangered with a population size of around 200. No viverrid species have been domesticated.
The thirty-three species of Viverridae are split into fourteen genera within four subfamilies : the three civet subfamilies Viverrinae , Hemigalinae , and Paradoxurinae , and the genet subfamily Genettinae . A fifth subfamily, Prionodontinae , was previously included in Viverridae, while the species in Genettinae were considered part of Viverrinae, but more recent genetic evidence resulted in the consensus to separate Prionodontinae into its own family and split out Genettinae into its own subfamily. Extinct species have also been placed into Viverrinae, as well as the extinct subfamily Lophocyoninae, though most extinct species have not been categorized into a subfamily. Nearly twenty extinct Viverridae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.
Conventions
Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species . Range maps are provided wherever possible; if a range map is not available, a description of the viverrid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.
Classification
The family Viverridae consists of 33 extant species belonging to 14 genera in 4 subfamilies and divided into dozens of extant subspecies. This does not include hybrid species or extinct prehistoric species.
Viverrids
The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis . This includes the promotion of the Prionodontinae subfamily into its own family, and the moving of the Poiana and Genetta genera out of the Viverrinae subfamily into their own Genettinae subfamily. There are several additional proposals which are disputed, such as splitting the golden palm civet into three species or adding three additional species to Genetta , which are not included here.[ 1] [ 2]
Subfamily Genettinae
Genus Genetta โ Cuvier , 1816 โ fourteen species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Abyssinian genet
G. abyssinica (Rรผppell , 1836)
Northeast Africa
Size : 40โ43 cm (16โ17 in) long, plus 38โ41 cm (15โ16 in) tail[ 3] Habitat : Forest, shrubland, and grassland[ 4] Diet : Primarily eats rodents, birds, and seeds, as well as insects and fruit[ 4] [ 5]
DD
Unknown [ 4]
Angolan genet
G. angolensis Bocage , 1882
Scattered south-central Africa
Size : 44โ48 cm (17โ19 in) long, plus 38โ43 cm (15โ17 in) tail[ 3] Habitat : Savanna[ 6] Diet : Believed to primarily eat insects, as well as fruit[ 6] [ 5]
LC
Unknown [ 6]
Aquatic genet
G. piscivora (Allen , 1919)
Central Africa
Size : 44โ50 cm (17โ20 in) long, plus 34โ42 cm (13โ17 in) tail[ 5] Habitat : Forest and inland wetlands[ 7] Diet : Primarily eats fish[ 7]
NT
10,000 [ 7]
Bourlon's genet
G. bourloni Gaubert , 2003
West Africa
Size : 49โ50 cm (19โ20 in) long, plus 40โ42 cm (16โ17 in) tail[ 5] Habitat : Forest[ 8] Diet : Unknown[ 8] [ 5]
VU
9,800 [ 8]
Cape genet
G. tigrina (Schreber , 1776)
G. t. methi
G. t. tigrina
South Africa
Size : 42โ58 cm (17โ23 in) long, plus 38โ46 cm (15โ18 in) tail[ 5] Habitat : Forest, shrubland, and grassland[ 9] Diet : Omnivorous; primarily eats insects and small mammals[ 9] [ 5]
LC
Unknown [ 9]
Common genet
G. genetta (Linnaeus , 1758)
G. g. afra
G. g. dongolana
G. g. felina
G. g. genetta
G. g. senegalensis
Bands of north, central, and south Africa and parts of Arabian peninsula (green), introduced in southwest Europe (red), formerly introduced in Nile river area (black)
Size : 46โ52 cm (18โ20 in) long, plus 42โ52 cm (17โ20 in) tail[ 3] Habitat : Forest, savanna, shrubland, and rocky areas[ 10] Diet : Primarily eats small mammals, as well as birds, other small vertebrates, insects, and fruit[ 10]
LC
Unknown [ 10]
Crested servaline genet
G. cristata Hayman , 1940
Northwest Africa
Size : 49โ63 cm (19โ25 in) long, plus 43 cm (17 in) tail[ 3] Habitat : Forest[ 11] Diet : Primarily eats insects, as well as small mammals, reptiles, and vegetables[ 11] [ 5]
VU
7,000 [ 11]
Giant forest genet
G. victoriae Thomas , 1902
Central Africa
Size : 55โ60 cm (22โ24 in) long, plus 41โ49 cm (16โ19 in) tail[ 5] Habitat : Forest[ 12] Diet : Primarily eats fruit, as well as rodents, birds, and insects[ 12] [ 5]
LC
Unknown [ 12]
Hausa genet
G. thierryi Matschie , 1902
West Africa
Size : 44โ45 cm (17โ18 in) long, plus 40โ43 cm (16โ17 in) tail[ 3] Habitat : Forest, savanna, and shrubland[ 13] Diet : Unknown[ 13] [ 5]
LC
Unknown [ 13]
Johnston's genet
G. johnstoni Pocock , 1908
Northwest Africa
Size : 47โ52 cm (19โ20 in) long, plus 46โ50 cm (18โ20 in) tail[ 5] Habitat : Forest, savanna, and shrubland[ 14] Diet : Believed to primarily eat insects[ 14] [ 5]
NT
Unknown [ 14]
King genet
G. poensis Waterhouse , 1838
Scattered parts of west Africa
Size : 42โ68 cm (17โ27 in) long, plus 35โ47 cm (14โ19 in) tail[ 5] Habitat : Forest[ 15] Diet : Unknown; believed to eat mammals and fruit[ 15]
DD
Unknown [ 15]
Pardine genet
G. pardina Geoffroy , 1832
Northwest Africa
Size : 41โ56 cm (16โ22 in) long, plus 39โ45 cm (15โ18 in) tail[ 5] Habitat : Forest, savanna, and shrubland[ 16] Diet : Primarily eats rodents, as well as insects, fruit, birds, and other small mammals[ 16] [ 5]
LC
Unknown [ 16]
Rusty-spotted genet
G. maculata (Gray , 1830)
Central and south Africa
Size : 41โ53 cm (16โ21 in) long, plus 39โ54 cm (15โ21 in) tail[ 3] Habitat : Forest, savanna, and shrubland[ 17] Diet : Primarily eats invertebrates, fish, amphibians, reptiles, small birds, eggs, and small mammals, as well as fruit, seeds, and berries[ 17] [ 5]
LC
Unknown [ 17]
Servaline genet
G. servalina Pucheran , 1855
Central africa
Size : 44โ51 cm (17โ20 in) long, plus 36โ49 cm (14โ19 in) tail[ 5] Habitat : Forest and shrubland[ 18] Diet : Primarily eats small mammals and insects, as well as snakes, birds, and fruit[ 18] [ 5]
LC
Unknown [ 18]
Genus Poiana โ Gray , 1865 โ two species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Central African oyan
P. richardsonii (Thomson , 1842)
P. r. ochracea
P. r. richardsonii
Central Africa
Size : 33โ38 cm (13โ15 in) long, plus 35โ40 cm (14โ16 in) tail[ 19] Habitat : Forest[ 20] Diet : Believed to eat small vertebrates and invertebrates[ 20] [ 3]
LC
Unknown [ 20]
West African oyan
P. leightoni Pocock , 1908
West Africa
Size : 30โ38 cm (12โ15 in) long, plus 35โ40 cm (14โ16 in) tail[ 3] Habitat : Forest[ 21] Diet : Believed to eat small vertebrates and invertebrates[ 21] [ 3]
VU
6,700-10,000 [ 21]
Subfamily Hemigalinae
Genus Chrotogale โ Thomas , 1912 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Owston's palm civet
C. owstoni Thomas , 1912
Southeast Asia around Vietnam
Size : 51โ63 cm (20โ25 in) long, plus 38โ48 cm (15โ19 in) tail[ 22] Habitat : Forest and shrubland[ 23] Diet : Believed to primarily eat earthworms and other invertebrates[ 23]
EN
Unknown [ 23]
Genus Cynogale โ Gray , 1837 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Otter civet
C. bennettii (Gray , 1837)
C. b. bennettii
C. b. lowei
Southeast Asia
Size : 57โ68 cm (22โ27 in) long, plus 12โ21 cm (5โ8 in) tail[ 22] Habitat : Forest and inland wetlands[ 24] Diet : Primarily eats fish, crabs, molluscs, small mammals, and birds[ 24]
EN
2,500 [ 24]
Genus Diplogale โ Thomas , 1912 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Hose's palm civet
D. hosei Thomas , 1892
Borneo in Southeast Asia
Size : 47โ54 cm (19โ21 in) long, plus 29โ34 cm (11โ13 in) tail[ 25] Habitat : Forest[ 26] Diet : Believed to primarily eat small fish, shrimp, crabs, and frogs as well as insects[ 26]
VU
9,500 [ 26]
Genus Hemigalus โ Jourdan , 1837 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Banded palm civet
H. derbyanus (Gray , 1837)
H. d. boiei
H. d. derbyanus
H. d. minor
H. d. sipora
Southeast Asia
Size : 45โ56 cm (18โ22 in) long, plus 25โ36 cm (10โ14 in) tail[ 22] Habitat : Forest[ 27] Diet : Primarily eats insects[ 27]
NT
Unknown [ 27]
Subfamily Paradoxurinae
Genus Arctictis โ Temminck , 1824 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Binturong
A. binturong (Raffles , 1821)
A. b. albifrons
A. b. binturong
A. b. kerkhoveni
A. b. menglaensis
A. b. penicillatus
A. b. whitei (Palawan binturong )
Southeast Asia
Size : 61โ96 cm (24โ38 in) long, plus 56โ89 cm (22โ35 in) tail[ 28] Habitat : Forest[ 29] Diet : Primarily eats fruit[ 29]
VU
Unknown [ 29]
Genus Arctogalidia โ Merriam , 1897 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Small-toothed palm civet
A. trivirgata (Gray , 1832)
A. t. bancana
A. t. fusca
A. t. inornata
A. t. leucotis
A. t. macra
A. t. major
A. t. millsi
A. t. minor
A. t. simplex
A. t. stigmaticus
A. t. sumatrana
A. t. tingia
A. t. trilineata
A. t. trivirgata
Southeast Asia
Size : 44โ60 cm (17โ24 in) long, plus 48โ66 cm (19โ26 in) tail[ 30] Habitat : Forest[ 31] Diet : Omnivorous; primarily eats fruit[ 31]
LC
Unknown [ 31]
Genus Macrogalidia โ Schwarz , 1910 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Sulawesi palm civet
M. musschenbroekii (Schlegel , 1877)
Sulawesi island in Southeast Asia
Size : 65โ72 cm (26โ28 in) long, plus 44โ54 cm (17โ21 in) tail[ 3] Habitat : Forest, shrubland, and grassland[ 32] Diet : Primarily eats rodents and palm fruit , as well as other small mammals, birds, fruit, and grass[ 32]
VU
9,000 [ 32]
Genus Paguma โ Gray , 1831 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Masked palm civet
P. larvata (H. Smith , 1827)
P. l. chichingensis
P. l. grayi
P. l. hainana
P. l. intrudens
P. l. janetta
P. l. jourdanii
P. l. lanigera
P. l. larvata
P. l. leucomystax
P. l. neglecta
P. l. nigriceps
P. l. ogilbyi
P. l. robusta
P. l. taivana
P. l. tytlerii
P. l. wroughtoni
East and southeast Asia
Size : 50โ76 cm (20โ30 in) long, plus 50โ64 cm (20โ25 in) tail[ 33] Habitat : Forest and shrubland[ 34] Diet : Omnivorous; primarily eats fruit[ 34]
LC
Unknown [ 34]
Genus Paradoxurus โ F. Cuvier , 1821 โ three species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Asian palm civet
P. hermaphroditus (Pallas , 1777)
P. h. balicus
P. h. bondar
P. h. canescens
P. h. canus
P. h. cochinensis
P. h. dongfangensis
P. h. enganus
P. h. exitus
P. h. hermaphroditus
P. h. javanica
P. h. kangeanus
P. h. laotum
P. h. lignicolor
P. h. milleri
P. h. minor
P. h. musanga
P. h. nictitans
P. h. pallasii
P. h. pallens
P. h. parvus
P. h. philippinensis
P. h. pugnax
P. h. pulcher
P. h. sacer
P. h. scindiae
P. h. senex
P. h. setosus
P. h. simplex
P. h. sumbanus
P. h. vellerosus
South and southeast Asia (green), introduced (red)
Size : 47โ57 cm (19โ22 in) long, plus 47โ56 cm (19โ22 in) tail[ 35] Habitat : Forest, shrubland, and grassland[ 36] Diet : Primarily eats rats and fruit, as well as insects and molluscs[ 36]
LC
Unknown [ 36]
Brown palm civet
P. jerdoni Blanford , 1885
P. j. caniscus
P. j. jerdoni
Southwest India
Size : 43โ62 cm (17โ24 in) long, plus 38โ53 cm (15โ21 in) tail[ 37] Habitat : Forest[ 38] Diet : Primarily eats fruit, as well as birds, rodents, and insects[ 38]
LC
Unknown [ 38]
Golden palm civet
P. zeylonensis (Schreber , 1778)
Sri Lanka
Size : 50โ58 cm (20โ23 in) long, plus 43โ53 cm (17โ21 in) tail[ 3] Habitat : Forest and shrubland[ 39] Diet : Primarily eats berries, fruits, and invertebrates, as well as small vertebrates[ 39]
LC
Unknown [ 39]
Subfamily Viverrinae
Genus Civettictis โ Pocock , 1915 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
African civet
C. civetta (Schreber , 1776)
C. c. australis
C. c. civetta
C. c. congica
C. c. pauli
C. c. schwarzi
C. c. volkmanni
Central and south Africa
Size : 60โ92 cm (24โ36 in) long, plus 43โ61 cm (17โ24 in) tail[ 40] Habitat : Forest, savanna, shrubland, and inland wetlands[ 41] Diet : Omnivorous; primarily eats fruit[ 41]
LC
Unknown [ 41]
Genus Viverra โ Linnaeus , 1758 โ four species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Large-spotted civet
V. megaspila Blyth , 1862
Southeast Asia
Size : 77โ90 cm (30โ35 in) long, plus 32โ40 cm (13โ16 in) tail[ 35] Habitat : Forest, shrubland, and inland wetlands[ 42] Diet : Primarily eats birds, frogs, snakes, small mammals, eggs, crabs, fish, fruit, and roots[ 35] [ 42]
EN
Unknown [ 42]
Large Indian civet
V. zibetha Linnaeus , 1758
V. z. ashtoni
V. z. hainana
V. z. picta
V. z. pruinosus
V. z. zibetha
Southeast Asia
Size : 50โ95 cm (20โ37 in) long, plus 38โ59 cm (15โ23 in) tail[ 35] Habitat : Forest and shrubland[ 43] Diet : Primarily eats fish, birds, lizards, frogs, insects, arthropods, and crabs, as well as poultry and rubbish[ 43]
LC
Unknown [ 43]
Malabar large-spotted civet
V. civettina Blyth , 1862
Southwest India
Size : 76 cm (30 in) long, plus 33 cm (13 in) tail[ 44] Habitat : Forest and inland wetlands[ 45] Diet : Unknown[ 45]
CR
200 [ 45]
Malayan civet
V. tangalunga Gray , 1832
V. t. lankavensis
V. t. tangalunga
Southeast Asia
Size : 58โ95 cm (23โ37 in) long, plus 30โ49 cm (12โ19 in) tail[ 46] Habitat : Forest and shrubland[ 47] Diet : Omnivorous[ 47]
LC
Unknown [ 47]
Genus Viverricula โ Hodgson , 1838 โ one species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Small Indian civet
V. indica (Desmarest , 1804)
V. i. atchinensis
V. i. baliensis
V. i. baptistae
V. i. deserti
V. i. indica
V. i. klossi
V. i. mayori
V. i. muriavensis
V. i. pallida
V. i. schlegelii
V. i. thai
V. i. wellsi
South and southeast Asia
Size : 45โ63 cm (18โ25 in) long, plus 30โ43 cm (12โ17 in) tail[ 48] Habitat : Forest, savanna, shrubland, grassland, and inland wetlands[ 49] Diet : Primarily eats rodents, birds, snakes, fruit, roots, carrion, and insects[ 49]
LC
Unknown [ 49]
References
^ Groves, C. P.; Rajapaksha, C.; Manemandra-Arachchi, K. (2009). "The taxonomy of the endemic golden palm civet of Sri Lanka" (PDF) . Zoological Journal of the Linnean Society . 155 : 238โ 251. doi :10.1111/j.1096-3642.2008.00451.x .
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^ a b c Gaubert, P.; Dinets, V.; Do Linh San, E. (2016). "Genetta victoriae " . IUCN Red List of Threatened Species . 2016 : e.T41703A45219531. doi :10.2305/IUCN.UK.2016-1.RLTS.T41703A45219531.en .
^ a b c Gaubert, P.; Do Linh San, E. (2015). "Genetta thierryi " . IUCN Red List of Threatened Species . 2015 : e.T41701A45219325. doi :10.2305/IUCN.UK.2015-4.RLTS.T41701A45219325.en .
^ a b c Gaubert, P.; Do Linh San, E. (2016). "Genetta johnstoni " . IUCN Red List of Threatened Species . 2016 : e.T8997A45198265. doi :10.2305/IUCN.UK.2016-1.RLTS.T8997A45198265.en .
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