List of ursids

IUCN Red List categories
Conservation status
EX	Extinct (0 species)
EW	Extinct in the wild (0 species)
CR	Critically Endangered (0 species)
EN	Endangered (0 species)
VU	Vulnerable (6 species)
NT	Near threatened (0 species)
LC	Least concern (2 species)

Brown bear in grass — Brown bear (*Ursus arctos*)

Ursidae is a family of mammals in the order Carnivora, which includes the giant panda, brown bear, and polar bear, and many other extant or extinct mammals. A member of this family is called a bear or an ursid. They are widespread across the Americas and Eurasia. Bear habitats are generally forests, though some species can be found in grassland and savana regions, and the polar bear lives in arctic and aquatic habitats. Most bears are 1.2–2 m (4–7 ft) long, plus a 3–20 cm (1–8 in) tail, though the polar bear is 2.2–2.44 m (7–8 ft) long, and some subspecies of brown bear can be up to 2.8 m (9 ft). Weights range greatly from the sun bear, which can be as low as 35 kg (77 lb), to the polar bear, which can be as high as 726 kg (1,600 lb). Population sizes vary, with six species classified as vulnerable with populations as low as 500, while the brown bear has a population of over 100,000 and the American black bear around 800,000. Many bear species primarily eat specific foods, such as seals for the polar bear or termites and fruit for the sloth bear, but with the exception of the giant panda, which exclusively eats bamboo, ursids are omnivorous when necessary. No ursid species have been domesticated, though some bears have been trained for entertainment.^[1]

The eight species of Ursidae are split into five genera in three subfamilies: the monotypic Ailuropodinae, the panda bears; Tremarctinae, the short-faced bears; and Ursinae, containing all other extant bears. Extinct species have also been placed into all three extant subfamilies, as well as three extinct ones: Agriotheriinae, Hemicyoninae, and Ursavinae. Over 100 extinct Ursidae species have been found, though due to ongoing research and discoveries the exact number and categorization is not fixed.

Conventions

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†". Population figures are rounded to the nearest hundred.

Classification

The family Ursidae consists of eight extant species belonging to five genera in three subfamilies and divided into dozens of extant subspecies. This does not include ursid hybrid species such as grizzly–polar bear hybrids or extinct prehistoric species.

Subfamily Ailuropodinae
- Genus Ailuropoda (panda bears): one species
Subfamily Tremarctinae
- Genus Tremarctos (short-faced bears): one species
Subfamily Ursinae
- Genus Helarctos: (sun bear): one species
- Genus Melursus: (sloth bear): one species
- Genus Ursus (bears): four species

Ursidae

Ursinae

	Ursus (American black / Asian black / brown / polar bear)

	Helarctos (sun bear)

Melursus (sloth bear)

Tremarctinae

Tremarctos (spectacled bear)

Ailuropodinae

Ailuropoda (giant panda)

Ursids

The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis; this includes the division of the giant panda into two subspecies. There are several additional proposals which are disputed, such as reclassifying the subspecies of the brown bear into a smaller set of clades,^[2]^[3] which are not included here.

Subfamily Ailuropodinae

Genus **Ailuropoda** – H. Milne-Edwards, 1870 – one species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Giant panda	A. melanoleuca (David, 1869) Two subspecies A. m. melanoleuca A. m. qinlingensis (Qinling panda)	Central China	Size: 150–180 cm (59–71 in) long, plus 10–15 cm (4–6 in) tail 80–123 kg (176–271 lb)^[4]^[5] Habitat: Forest^[6] Diet: Eats only bamboo^[6]	VU 500–1,000 ^[6]

Subfamily Tremarctinae

Genus **Tremarctos** – Gervais, 1855 – one species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Spectacled bear	T. ornatus (F. Cuvier, 1825)	Andes mountains in South America	Size: 120–200 cm (47–79 in) long, plus 7 cm (3 in) tail 60–175 kg (132–386 lb)^[7] Habitat: Shrubland, grassland, and forest^[8] Diet: Primarily eats bromeliads and palm trees, as well as cattle, other mammals, and fruit^[8]	VU 2,500–10,000 ^[8]

Subfamily Ursinae

Genus **Helarctos** – Horsfield, 1825 – one species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Sun bear	H. malayanus (Raffles, 1821) Two subspecies H. m. euryspilus (Bornean sun bear) H. m. malayanus (Malayan sun bear)	Southeast Asia (current range in brown, former in black)	Size: 120–150 cm (47–59 in) long, plus 3–7 cm (1–3 in) tail 35–80 kg (77–176 lb)^[9]^[10] Habitat: Forest and shrubland^[11] Diet: Primarily eats termites, ants, beetle larvae, bee larvae, honey, and fruit^[11]	VU 50,000^[12] ^[11]

Genus **Melursus** – Meyer, 1793 – one species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
Sloth bear	M. ursinus (Shaw, 1791) Two subspecies M. u. inornatus (Sri Lankan sloth bear) M. u. ursinus (Indian sloth bear)	India (current range in green, former in black)	Size: 150–180 cm (59–71 in) long, plus 7–12 cm (3–5 in) tail 54–141 kg (119–311 lb)^[13] Habitat: Shrubland, grassland, forest, and savanna^[14] Diet: Primarily eats termites and fruit^[14]	VU 6,000–20,000^[14]^[15] ^[14]

Genus **Ursus** – Linnaeus, 1758 – four species
Common name	Scientific name and subspecies	Range	Size and ecology	IUCN status and estimated population
American black bear	U. americanus Pallas, 1780 Sixteen subspecies U. a. altifrontalis (Olympic black bear) U. a. amblyceps (New Mexico black bear) U. a. americanus (Eastern black bear) U. a. californiensis (California black bear) U. a. carlottae (Haida Gwaii black bear) U. a. cinnamomum (Cinnamon bear) U. a. emmonsii (Glacier bear) U. a. eremicus (East Mexican black bear) U. a. floridanus (Florida black bear) U. a. hamiltoni (Newfoundland black bear) U. a. kermodei (Kermode bear) U. a. luteolus (Louisiana black bear) U. a. machetes (West Mexican black bear) U. a. perniger (Kenai black bear) U. a. pugnax (Dall Island black bear) U. a. vancouveri (Vancouver Island black bear)	North America (current range in red, former in pink)	Size: 120–200 cm (47–79 in) long, plus 8–14 cm (3–6 in) tail 39–409 kg (86–902 lb)^[16] Habitat: Forest, inland wetlands, grassland, shrubland, and desert^[17] Diet: Omnivorous; eats vegetation, roots, buds, fruit, nuts, insects, fish, mammals, and carrion^[17]	LC 735,000–941,000^[18] ^[17]
Asian black bear	U. thibetanus Cuvier, 1823 Seven subspecies U. t. formosanus (Formosan black bear) U. t. gedrosianus (Balochistan black bear) U. t. japonicus (Japanese black bear) U. t. laniger (Himalayan black bear) U. t. mupinensis (Indochinese black bear) U. t. thibetanus (Tibetan black bear) U. t. ussuricus (Ussuri black bear)	South and East Asia (current range in brown, former in black)	Size: 120–180 cm (47–71 in) long, plus 6–11 cm (2–4 in) tail 65–150 kg (143–331 lb)^[19] Habitat: Forest, inland wetlands, grassland, and shrubland^[20] Diet: Eats vegetation, insects, fruit, nuts, ungulates, and livestock^[20]	VU 50,000^[12] ^[20]
Brown bear	U. arctos Linnaeus, 1758 Sixteen subspecies U. a. alascensis (Alaskan grizzly bear) U. a. arctos (Eurasian brown bear) U. a. beringianus (Kamchatka brown bear) U. a. californicus (California grizzly bear) U. a. collaris (East Siberian brown bear) U. a. crowtheri (Atlas bear)† U. a. dalli (Dall Island brown bear) U. a. gyas (Alaska Peninsula brown bear) U. a. horribilis (Grizzly bear) U. a. isabellinus (Himalayan brown bear) U. a. lasiotus (Ussuri brown bear) U. a. middendorffi (Kodiak bear) U. a. pruinosus (Tibetan blue bear) U. a. sitkensis (ABC Islands bear) U. a. stikeenensis (Stickeen brown bear) U. a. syriacus (Syrian brown bear)	Northern North America and Europe, and northern and central Asia	Size: 100–280 cm (39–110 in) long, plus 6–20 cm (2–8 in) tail 80–550 kg (176–1,213 lb)^[21] Habitat: Desert, forest, inland wetlands, grassland, and shrubland^[22] Diet: Omnivorous; eats grasses, herbs, roots, berries, nuts, insects, mammals, and fish^[22]	LC 110,000 ^[22]
Polar bear	U. maritimus Mulgrave, 1774	Polar North America and Asia	Size: 220–244 cm (87–96 in) long, plus 7–13 cm (3–5 in) tail 408–726 kg (900–1,600 lb)^[23] Habitat: Marine oceanic, shrubland, forest, grassland, marine coastal/supratidal, and marine intertidal^[24] Diet: Primarily eats seals, as well as walruses, beluga whales, birds, fish, vegetation and kelp^[24]	VU 23,000^[25] ^[24]

Prehistoric ursids

In addition to extant bears, a number of prehistoric species have been discovered and classified as a part of Ursidae. In addition to being placed within the three extant subfamilies, they have been categorized within the extinct subfamilies Agriotheriinae, Hemicyoninae, and Ursavinae, some of which are subdivided into named tribes. There is no generally accepted classification of extinct ursid species. The species listed here are based on data from the Paleobiology Database, unless otherwise cited. Where available, the approximate time period the species was extant is given in millions of years before the present (Mya), also based on data from the Paleobiology Database.^[26] All listed species are extinct; where a genus or subfamily within Ursidae comprises only extinct species, it is indicated with a dagger symbol †.

Subfamily Agriotheriinae†
- Genus Agriotherium†
  - A. africanum (3.6–2.5 Mya)
  - A. gregoryi
  - A. inexpetans (12–5.3 Mya)
  - A. insigne
  - A. schneideri (14–2.5 Mya)
  - A. sivalensis (5.4–3.6 Mya)
Subfamily Ailuropodinae
- Tribe Ailuropodini
  - Genus Agriarctos†
  - Genus Ailurarctos†
    - A. lufengensis^[27]
  - Genus Ailuropoda
    - A. baconi (2.6–0.012 Mya)
    - A. fovealis
    - A. microta (2.6–0.78 Mya)
    - A. wulingshanensis (2.6–0.78 Mya)
  - Genus Kretzoiarctos† (12–8.7 Mya)
    - K. beatrix (12–8.7 Mya)
- Tribe Indarctini†
  - Genus Indarctos†
    - I. anthracitis
    - I. arctoides (9.7–8.7 Mya)
    - I. atticus (8.7–5.3 Mya)
    - I. nevadensis (11–4.9 Mya)
    - I. oregonensis (11–4.9 Mya)
    - I. salmontanus
    - I. vireti
    - I. zdanskyi
  - Genus Miomaci† (12–9.7 Mya)
    - M. pannonicum (12–9.7 Mya)
Subfamily Hemicyoninae†
- Tribe Cephalogalini†
  - Genus Adelpharctos† (34–23 Mya)
    - A. ginsburgi (29–23 Mya)
    - A. mirus (34–28 Mya)
  - Genus Cephalogale†
    - C. geoffroyi
    - C. meschethense (29–23 Mya)
    - C. minor (34–28 Mya)
  - Genus Cyonarctos† (29–23 Mya)
    - C. dessei (29–23 Mya)
  - Genus Filholictis†
    - F. filholi
  - Genus Phoberogale†
    - P. depereti
    - P. shareri (31–20 Mya)
- Tribe Hemicyonini†
  - Genus Dinocyon†
    - D. aurelianensis
    - D. sansaniensis
    - D. thenardi (17–15 Mya)
  - Genus Hemicyon†
    - H. barbouri (14–10 Mya)
    - H. goriachensis
    - H. grivensis
    - H. minor
    - H. sansaniensis (16–12 Mya)
    - H. statzlingii
  - Genus Zaragocyon† (23–20 Mya)
    - Z. daamsi (23–20 Mya)
- Tribe Phoberocyonini†
  - Genus Phoberocyon†
    - P. aurelianensis (21–7.2 Mya)
    - P. huerzeleri
    - P. johnhenryi (21–15 Mya)
  - Genus Plithocyon† (16–11 Mya)
    - P. armagnacensis (16–11 Mya)
    - P. barstowensis (16–13 Mya)
    - P. ursinus (16–13 Mya)
Subfamily Tremarctinae
- Genus Arctodus (short-faced bear)† (2.2–0.012 Mya)
  - A. pristinus (lesser short-faced bear) (2.2–0.3 Mya)
  - A. simus (giant short-faced bear) (1.1–0.012 Mya)
- Genus Arctotherium†
  - A. angustidens (1.2–0.7 Mya)
  - A. bonariense (0.6–0.037 Mya)
  - A. tarijense (0.5–0.010 Mya)
  - A. vetustum (0.7–0.3 Mya)
  - A. wingei (0.1–0.012 Mya)
- Genus Plionarctos† (10.3–1.8 Mya)
  - P. edensis (10.3–4.9 Mya)
  - P. harroldorum (4.9–1.8 Mya)
- Genus Tremarctos (1.8 Mya–present)
  - T. floridanus (Florida spectacled bear) (1.8–0.012 Mya)
Subfamily Ursavinae†
- Tribe Ursavini†
  - Genus Ursavus†
    - U. brevirhinus (16–9.7 Mya)
    - U. elmensis (dawn bear) (16–13 Mya)
    - U. pawniensis (24–5.3 Mya)
    - U. primaevus (14–9.7 Mya)
Subfamily Ursinae
- Genus Aurorarctos† (15–12.5 Mya)
  - A. tirawa (15–12.5 Mya)
- Genus Protarctos† (4.9–3.6 Mya)
  - P. abstrusus (4.9–3.6 Mya)
- Genus Ursus
  - U. arctos priscus (steppe brown bear)^[a]^[28]
  - U. arvernensis (3.2–2.5 Mya)
  - U. deningeri (0.79–0.12 Mya) (Deninger's bear)^[29]
  - U. dolinensis (Gran Dolina bear)^[30]
  - U. dentrificius (2.6–0.12 Mya)
  - U. etruscus (Etruscan bear)^[31]
  - U. eogroenlandicus
  - U. ingressus (Gamssulzen cave bear)^[32]
  - U. inopinatus
  - U. jenaensis
  - U. labradorensis
  - U. malayanus (0.13–0.012 Mya)
  - U. maritimus tyrannus (tyrant polar bear)^[a]^[33]
  - U. minimus (Auvergne bear) (4.2–2.5 Mya)
  - U. optimus
  - U. praemalayanus
  - U. priscus
  - U. rossicus (pleistocene small cave bear)^[34]
  - U. spelaeus (cave bear) (2.6–0.012 Mya)
  - U. spitzbergensis
  - U. vitabilis

Restoration of Arctodus simus
Restoration of Arctotherium bonariense
Restoration of Ursus spelaeus (cave bear)
Prehistoric art of Ursus spelaeus (cave bear)

Notes

^ ^a ^b Extinct prehistoric subspecies of an extant species

References

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[extinct-subspecies-28] Extinct prehistoric subspecies of an extant species

[entertainment-1] Findeizen, Nikolai (2008). History of Music in Russia from Antiquity to 1800, Vol. 1: From Antiquity to the Beginning of the Eighteenth Century. Indiana University Press. p. 201. ISBN 978-0-253-02637-8.

[BrownBearSubspecies1-2] Calvignac, S.; Hughes, S.; Tougard, C.; Michaux, J.; Thevenot, M.; Philippe, M.; Hamdine, W.; Hanni, C. (2008). "Ancient DNA evidence for the loss of a highly divergent brown bear clade during historical times" (PDF). Molecular Ecology. 17 (8): 1962–1970. doi:10.1111/j.1365-294x.2008.03631.x. PMID 18363668. S2CID 23361337.

[BrownBearSubspecies2-3] Lan, T.; Gill, S.; Bellemain, E.; Bischof, R.; Zawaz, M. A.; Lindqvist, C. (2017). "Evolutionary history of enigmatic bears in the Tibetan Plateau–Himalaya region and the identity of the yeti". Proceedings of the Royal Society B: Biological Sciences. 284 (1868): 20171804. doi:10.1098/rspb.2017.1804. PMC 5740279. PMID 29187630.

[ADWGiantpanda-4] Bies, LeeAnn (2002). "Ailuropoda melanoleuca". Animal Diversity Web. University of Michigan. Retrieved March 24, 2020.

[Giantpandatail-5] "Physical Description". Knowledge Hub. World Wide Fund for Nature. Retrieved March 24, 2020.

[IUCNGiantpanda-6] Swaisgood, R.; Wang, D.; Wei, F. (2017) [errata version of 2016 assessment]. "Ailuropoda melanoleuca". IUCN Red List of Threatened Species. 2016: e.T712A121745669.

[Spectacledbearsize-7] "Spectacled bear (Tremarctos ornatus)". ARKive. Wildscreen. Archived from the original on June 14, 2017. Retrieved March 25, 2020.

[IUCNSpectacledbear-8] Velez-Liendo, X.; García-Rangel, S. (2018) [errata version of 2017 assessment]. "Tremarctos ornatus". IUCN Red List of Threatened Species. 2017: e.T22066A45034047. doi:10.2305/IUCN.UK.2017-3.RLTS.T22066A45034047.en.

[Sunbearsize-9] "Malayan sun bear (Helarctos malayanus)". ARKive. Wildscreen. Archived from the original on December 21, 2014. Retrieved March 26, 2020.

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[Ursusdolinensis-31] Garcı́a, N.; Arsuaga, J. L. (June 15, 2001). "Ursus dolinensis4: a new species of Early Pleistocene ursid from Trinchera Dolina, Atapuerca (Spain)". Comptes Rendus de l'Académie des Sciences, Série IIA. 332 (11): 717–725. Bibcode:2001CRASE.332..717G. doi:10.1016/S1251-8050(01)01588-9.

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List of ursids

Conventions

Classification

Ursids

Subfamily Ailuropodinae

Subfamily Tremarctinae

Subfamily Ursinae

Prehistoric ursids

See also

Notes

References

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